

Natural Resources
Conservation Service
Ecological site R058CY070ND
Badland Fan
Last updated: 4/21/2025
Accessed: 05/19/2025
General information
Provisional. A provisional ecological site description has undergone quality control and quality assurance review. It contains a working state and transition model and enough information to identify the ecological site.
MLRA notes
Major Land Resource Area (MLRA): 058C–Northern Rolling High Plains, Northeastern Part
MLRA 58C covers 2,320 square miles and encompasses approximately 1.8 million acres. MLRA 58C spans two states, with 96 percent located in North Dakota and the remaining 4 percent is in Montana. The MLRA 58C landscape is characterized by steeply sloping dissected badlands along the Little Missouri River and its tributaries. Primary land uses are of rangeland for grazing and wildlife habitat. Microclimates inherent in badlands landscapes influence both variety and abundance of vegetation in MLRA 58C. South- and west-facing exposures are dry, hot, and sparsely vegetated. More humid and cooler north- and east-facing exposures are favorable for abundant forage and woody vegetation.
MLRA 58C is known as the Little Missouri Badlands, which formed when the Little Missouri River was diverted along a shorter, steeper course by Pleistocene glaciers. Due to the resulting increased gradient after its eastward diversion by the glaciers, the Little Missouri River began rapidly downcutting into the soft, calcareous sedimentary shale, siltstone, and sandstone of the Fort Union and Hell Creek geological formations. This rapid downcutting eroded and carved the badlands of the MLRA. This cycle of erosion and deposition continues today.
Most of the soils in MLRA 58C developed from residuum weathered in place. As a result of constant erosion and deposition, the majority of soils in MLRA 58C are Entisols and Inceptisols. Mollisols formed on the high, stable drainageway divides and plateaus above the steeper, dissected hillslopes and fans that define the Little Missouri Badlands. Elevation ranges from 1,835 feet (560 meters) to 3,400 feet (1,036 meters). The Little Missouri River flows through the entire length of MLRA 58C and empties into Lake Sakakawea that was formed by the Garrison Dam on the Missouri River.
Classification relationships
Level IV Ecoregions of the Conterminous United States: 43b-Little Missouri Badlands.
Ecological site concept
The Badland Fan ecological site is on depositional fans at the base of steep badland escarpments in MLRA 58C. Rapid, unimpeded overland flow of water during rainfall events causes erosion of the steep, sparsely vegetated or non-vegetated badland escarpments and deposition of sediments onto the alluvial fans below them. When the sedimentary parent material of the escarpment is sodic, this site often occurs in conjunction with the Thin Claypan site. Slopes typically range from 0 to 25 percent; some gullied areas with steeper slopes also occur.
Soils on this site are well drained and very deep with soft, sedimentary bedrock parent material deeper than 60 inches. Typically, a thin A horizon is underlain by numerous stratified layers of alluvial sediments. In many cases, a transitional AC horizon develops directly below the surface A horizon. Surface textures are usually loam or silt loam that will form a ribbon 1 to 2 inches long. Carbonates are throughout the soil profile. The C horizon is typically thinly stratified layers of fine sandy loam to silty clay loam. These thin, stratified layers throughout the C horizons are generally less than 2 inches thick and form as the result of erosion of the steep badland escarpment and constant deposition of sediments onto the alluvial fan at the base of the escarpment.
On the alluvial fan directly below the badland escarpment, vegetation is usually relatively sparse and bare ground is visibly noticeable since frequent erosion and deposition make it difficult for plants to establish. Vegetation and production gradually improve as the site stabilizes farther away from the base of the escarpment. The central concept of the soil chemistry of this site is non-saline to slightly saline. However, currently included in this site are some soils with E.C. >8 dS/m (see Site Development and Testing Plan). Carbonates at the surface and sparse vegetation on Badland Fan sites cause the soils on this site to be susceptible to both wind and water erosion. The hazard of water erosion greatly increases on slopes greater than 6 percent.
Badland, Shallow Loamy, and Very Shallow ecological sites occur higher on the landscape while Limy Residual sites occur lower. Thin Claypan sites occur on similar landscape positions as the Badland Fan ecological site; vegetation on the Thin Claypan site is strongly impacted by dense soil structure and/or high sodium content with accumulated salts.
To see a full copy of the ecological site description with all tables and the full version 5 rangeland health worksheet, please use the following hyperlink:
https://efotg.sc.egov.usda.gov/references/public/ND/58C_Badland_Fan_Narrative_Final_Ref_FSG.pdf
Associated sites
R058CY103ND |
Badland This site is on the very steep, sparsely vegetated badland escarpment above the Badland Fan site. The Badland site is characterized by exposed, soft, sedimentary siltstone and shale bedrock that is actively and constantly eroding. |
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R058CY081ND |
Thin Claypan This site is on similar landscape positions as the Badland Fan site; the sites may be in complex on long, linear alluvial fans below sodium-affected remnants of badland escarpments. Soft sedimentary bedrock parent material that is eroding off the badland remnant is sodium-affected, so soils that develop on the long, alluvial fan below are also high in sodium. Thin Claypan soils on these depositional fans have not developed the characteristic dense claypan layer since they are subjected to constant erosion and deposition. The sparse vegetation and the plant communities reflect the presence of elevated sodium in the soil. |
R058CY086ND |
Shallow Loamy This site is higher on the landscape than the Badland Fan ecological site. The soils have soft, sedimentary bedrock at a depth of 10 to 20 inches; this bedrock affects root growth. |
R058CY083ND |
Very Shallow This site occurs on ridges. The soils are either <10 inches to soft, sedimentary bedrock or are <20 inches to porcelanite (Kirby soils). Root growth and available water capacity are affected. |
R058CY079ND |
Limy Residual This site occurs on relatively stable, long, linear alluvial fans below the Badland Fan site. As the front of the alluvial fan extends farther from the eroding badland escarpment, it receives progressively less deposition. Although still a relatively young landform, the alluvial fan in this position receives less deposition from the badland escarpment; is more stable; and vegetative cover improves. Consequently, the soils on the Limy Residual ecological site are better developed than soils on the Badland Fan site. The surface horizon is thin on Limy Residual sites; these soils generally do not have a mollic epipedon. Soils on the Limy Residual site are very deep, medium textured soils that are calcareous within 8 inches of the soil surface. They have similar textures to Badland Fan soils. The Limy Residual site has better vegetative production than the Badland Fan site. |
Similar sites
R058CY081ND |
Thin Claypan This site is on similar landscape positions as the Badland Fan site; the sites may be in complex on long, linear alluvial fans below sodium-affected remnants of badland escarpments. Soft sedimentary bedrock parent material that is eroding off the badland remnant is sodium-affected, so soils that develop on the long, alluvial fan below are also high in sodium. Thin Claypan soils on these depositional fans have not developed the characteristic dense claypan layer since they are subjected to constant erosion and deposition. The sparse vegetation and the plant communities reflect the presence of elevated sodium in the soil. |
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R058CY079ND |
Limy Residual This site occurs on relatively stable, long, linear alluvial fans below the Badland Fan site. As the front of the alluvial fan extends farther from the eroding badland escarpment, it receives progressively less deposition. Although still a relatively young landform, the alluvial fan in this position receives less deposition from the badland escarpment; is more stable; and vegetative cover improves. Consequently, the soils on the Limy Residual ecological site are better developed than soils on the Badland Fan site. The surface horizon is thin on Limy Residual sites; these soils generally do not have a mollic epipedon. Soils on the Limy Residual site are very deep, medium textured soils that are calcareous within 8 inches of the soil surface. They have similar textures to Badland Fan soils. The Limy Residual site has better vegetative production than the Badland Fan site. |

Figure 1. Badland Fan ecological site in relationship to Badland ecological site

Figure 2. Badland Fan ecological site in relation to Badland ecological site. Plant Community Phases 1.1 and 2.1 are stabilized, while Plant Community Phases 1.2 and 2.2 have active erosion/deposition.
Table 1. Dominant plant species
Tree |
Not specified |
---|---|
Shrub |
Not specified |
Herbaceous |
(1) Pascopyrum smithii |
Physiographic features
This site occurs on nearly level to moderately steep depositional alluvial fans at the base of steep badland escarpments in MLRA 58C. Gullied areas also occur.
Table 2. Representative physiographic features
Landforms |
(1)
Alluvial fan
|
---|---|
Runoff class | Low to very high |
Flooding frequency | None |
Ponding frequency | None |
Elevation | 1,835 – 3,400 ft |
Slope | 2 – 25% |
Water table depth | 60 – 80 in |
Aspect | Aspect is not a significant factor |
Climatic features
MLRA 58C is considered to have a continental climate with cold winters and hot summers, low humidity, light rainfall, and much sunshine. Extremes in temperature are common and characteristic of the MLRA. The continental climate is the result of the location of this MLRA in the geographic center of North America. There are few natural barriers on the northern Great Plains, so air masses move unobstructed across the plains and account for rapid changes in temperature.
Annual precipitation ranges from 14 to 17 inches per year. The normal average annual temperature is about 41° F. January is the coldest month with an average temperature of about 17° F. July is the warmest month with an average temperature of about 70° F. The range of normal average monthly temperatures between the coldest and warmest months is 53° F. This large temperature range attests to the continental nature of the MLRA 58C climate. Wind speeds average about 11 miles per hour, ranging from about 13 miles per hour during the spring to about 10 miles per hour during the summer. Daytime wind speeds are generally stronger than nighttime wind speeds, and occasional strong storms may bring brief periods of high winds with gusts to more than 50 miles per hour.
Growth of native cool-season plants begins in late March and continues to early to mid-July. Native warm-season plants begin growth in mid-May and continue to the end of August. Greening up of cool-season plants can occur in September and October when adequate soil moisture is present.
Table 3. Representative climatic features
Frost-free period (characteristic range) | 91-100 days |
---|---|
Freeze-free period (characteristic range) | 119-123 days |
Precipitation total (characteristic range) | 15-16 in |
Frost-free period (actual range) | 84-102 days |
Freeze-free period (actual range) | 116-123 days |
Precipitation total (actual range) | 14-16 in |
Frost-free period (average) | 95 days |
Freeze-free period (average) | 121 days |
Precipitation total (average) | 15 in |
Figure 3. Monthly precipitation range
Figure 4. Monthly minimum temperature range
Figure 5. Monthly maximum temperature range
Figure 6. Monthly average minimum and maximum temperature
Figure 7. Annual precipitation pattern
Figure 8. Annual average temperature pattern
Climate stations used
-
(1) WATFORD CITY 14S [USC00329246], Grassy Butte, ND
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(2) TROTTERS 3 SSE [USC00328812], Beach, ND
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(3) MEDORA 7 E [USW00094080], Fairfield, ND
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(4) AMIDON [USC00320209], Amidon, ND
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(5) CARLYLE 13 NW [USC00241518], Wibaux, MT
Influencing water features
Overland flow carrying sediments eroded from the adjacent badland escarpment deposits those sediments on the alluvial fan at the base of the escarpment. A seasonal water table is deeper than 5 feet throughout the growing season. Surface infiltration and permeability through the profile are moderately slow to moderate. Water loss is through evapotranspiration and percolation below the root zone.

Figure 9. Debris from concentrated flow where runoff potential is very high due to slope. Note sediment on soil surface from recent water flow event.
Soil features
Soils associated with the Badland Fan ecological sites are in the Entisol order and are classified further as Aridic Ustorthents. These soils were developed under prairie vegetation. They formed in calcareous, medium-textured slope alluvium eroding from the adjacent sparsely vegetated, very steep, soft sedimentary bedrock escarpment (i.e., Badland). The soils on this site are very deep; typically, the depth to soft sedimentary bedrock is more than 80 inches but may be as shallow as 60 inches in some soils. The soils are well drained.
The sediments deposited by overland water flow at the base of the very steep badland escarpments are recent alluvium originally formed from soft sedimentary bedrock laid down during the Tertiary and late Cretaceous periods. The soft sedimentary bedrock is primarily calcareous siltstone, mudstone, or shale. Textures of the A horizon (i.e., surface layer) are typically loam or silt loam but can include fine sandy loam, clay loam, or silty clay loam. The A horizon is light brownish gray in color and typically ranges from about 0.5 to 3 inches thick. Typically, a thin A horizon is underlain by numerous stratified layers of alluvial sediments. In some cases, a transitional AC horizon (i.e., subsurface layer) has developed directly below the A horizon.
Substratum textures range from fine sandy loam to silty clay loam but are commonly loam or clay loam and will form a ribbon less than 2 inches long. Typically, these C horizon(s) are made up of numerous thin layers of various soil textures and colors. Buried A horizons can occur. Some strata may contain small fragments and chips of scoria if the adjacent escarpment is capped with porcelanite.
Soil reaction typically is slightly alkaline or moderately alkaline (pH 7.4 to 8.4) in the surface layer and ranges to strongly alkaline (pH 8.5 to 9.0) in the substratum. Typically, soil salinity is none to very slight (E. C. <4 dS/m); however slight salinity (E.C. 4 to 8 dS/m) is allowable. Sodicity is typically none to moderate (SAR 0 to 10); however, is some soils it ranges to high (SAR 10 to 20). Calcium carbonate content typically ranges from low to high (5 to 30 percent).
Soil on the Badland Fan ecological site is susceptible to water erosion. The hazard of water erosion increases where vegetative cover is sparse or non-existent.
The major soil series which characterizes the Badland Fan ecological site is Patent. Also currently included in this site is Sham; however, some Sham components have higher salinity levels than the central concept of this site.
Access Web Soil Survey (https://websoilsurvey.sc.egov.usda.gov/App/WebSoilSurvey.aspx) for specific local soils information.
Table 4. Representative soil features
Parent material |
(1)
Slope alluvium
–
shale and siltstone
(2) Slope alluvium – mudstone |
---|---|
Surface texture |
(1) Loam (2) Silt loam |
Family particle size |
(1) Loamy |
Drainage class | Well drained |
Permeability class | Moderately slow to moderate |
Depth to restrictive layer | 60 – 80 in |
Surface fragment cover <=3" | 5% |
Surface fragment cover >3" | Not specified |
Available water capacity (0-40in) |
5 – 9 in |
Calcium carbonate equivalent (0-40in) |
5 – 30% |
Electrical conductivity (0-40in) |
8 mmhos/cm |
Sodium adsorption ratio (0-40in) |
20 |
Soil reaction (1:1 water) (0-40in) |
7.4 – 9 |
Subsurface fragment volume <=3" (0-40in) |
24% |
Subsurface fragment volume >3" (0-40in) |
Not specified |
Ecological dynamics
The Badland Fan ecological site is on depositional fans at the base of steep badland escarpments in MLRA 58C. Rapid, unimpeded overland flow of water during rainfall events causes erosion of the steep, sparsely vegetated or non-vegetated badland escarpments and deposition of sediments onto the alluvial fans below them. As such, management of the Badland Fan ecological site does not change the dynamics of the erosion of the steep badland escarpment and constant deposition of sediments onto the alluvial fan at the base of the escarpment. Prescribed grazing may help stabilize the areas downslope of the depositional areas.
On the alluvial fan directly below the badland escarpment, vegetation is usually relatively sparse and bare ground is visibly noticeable since frequent erosion and deposition make it difficult for plants to establish. Vegetation and production gradually improve as the site stabilizes farther away from the base of the escarpment. As water flow patterns change, areas may be stabilized while stabilized areas may begin to erode. Carbonates at the surface and sparse vegetation on Badland Fan sites cause the soils on this site to be susceptible to both wind and water erosion.
Occasionally a rotational slide occurs, the sliding mass may stay intact and result in relatively small changes in site conditions. However, it may also break up into portions of different sizes and settle out in different orientations. Often, much of the vegetation present before the slide continues growing after being relocated by the slide. The result is a mosaic of various sized segments that can markedly differ in soil characteristics, aspect, areal extent, existing vegetation, and other factors. Over time, different ecological sites may develop, e.g., Badland Fan, Community Phase 2.2. As such, care should be given to identify any rotational slides, as well as whether the slide has resulted in a change in states, community phases, or perhaps ecological site(s).
The hazard of water erosion greatly increases on slopes greater than 6 percent and when the Badland Fan ecological site is in areas of concentrated flows below Badland ecological sites. Soil deposition not only occurs on soil surface but may also cover entire plants.
This ecological site description is based on nonequilibrium ecology and resilience theory and utilizes a State-and-Transition Model (STM) diagram to organize and communicate information about ecosystem change as a basis for management. The ecological dynamics characterized by the STM diagram reflect how changes in ecological drivers, feedback mechanisms, and controlling variables can maintain or induce changes in plant community composition (phases and/or states). The application of various management actions, combined with weather variables, impact the ecological processes which influence the competitive interactions, thereby maintaining or altering plant community structure.
Prior to European influence, the historical disturbance regime for MLRA 58C included frequent fires, both anthropogenic and natural in origin. Most fires, however, were anthropogenic fires set by Native Americans. Native Americans set fires in all months except perhaps January. These fires occurred in two peak periods, one from March-May with the peak in April and another from July-November with the peak occurring in October. Most of these fires were scattered and of small extent and duration. The grazing history would have involved grazing and browsing by large herbivores (such as American bison, elk, pronghorn, mule deer, and whitetail deer). Herbivory by small mammals, insects, nematodes, and other invertebrates are also important factors influencing the production and composition of the communities. Grazing and fire interaction, particularly when coupled with drought events, influenced the dynamics discussed and displayed in the following state and transition diagram and descriptions.
Following European influence, this ecological site generally has had a history of grazing by domestic livestock, particularly cattle, which along with other related activities (e.g., fencing, water development, fire suppression) has changed the disturbance regime of the site. Changes will occur in the plant communities due to these and other factors.
Weather fluctuations, coupled with managerial factors, may lead to changes in the plant communities and may, under adverse impacts, result in a slow decline in vegetative vigor and composition. However, under favorable conditions the botanical composition may resemble that prior to European influence.
Five vegetative states have been identified for the site (Reference, Native/Invaded, Invaded, Invaded Conifer, and Go-Back). Within each state, one or more community phases have been identified. These community phases are named based on the more dominant and visually conspicuous species and have been determined by study of historical documents, relict areas, scientific studies, and ecological aspects of plant species and plant communities. Transitional pathways and thresholds have been determined through similar methods.
State 1: Reference State represents the natural range of variability that dominated the dynamics of this ecological site prior to European influence. Dynamics of the state were largely determined by variations in climate and weather (e.g., drought), as well as that of fire (e.g., timing, frequency) and grazing by native herbivores (e.g., frequency, intensity, selectivity). Due to those variations, the Reference State is thought to have shifted temporally and spatially between three community phases.
Currently the primary disturbances include widespread introduction of exotic plants, concentrated livestock grazing, lack of fire, and perhaps long-term non-use or very light grazing, and no fire. Because of these changes, particularly the widespread occurrence of exotic species, as well as other environmental changes, the Reference State is considered to no longer exist. Thus, the presence of exotic plants on the site precludes it from being placed in the Reference State. It must then be placed in one of the other states, commonly State 2: Native/Invaded State (T1A).
State 2: Native/Invaded State. Colonization of the site by exotic plants results in a transition from State 1: Reference State to State 2: Native/Invaded State (T1A). This transition was inevitable and often resulted from colonization by exotic cool-season grasses such as Kentucky bluegrass, crested wheatgrass, and/or annual bromes, which have been particularly and consistently invasive under extended periods of no use and no fire. Other exotic plants (e.g., Canada thistle, hound’s tongue) are also known to invade the site.
Three community phases have been identified for this state and are similar to the community phases in the Reference State but have now been invaded by exotic cool-season grasses. These exotic cool- season grasses can be expected to increase. As that increase occurs, plants more desirable to wildlife and livestock decline. A decline in forb diversity can also be expected. Under non-use or minimal use management, mulch increases and may become a physical barrier to plant growth. This also changes the micro-climate near the soil surface and may alter infiltration, nutrient cycling, and biological activity near the soil surface. As a result, these factors combined with shading cause desirable native plants to have increasing difficulty remaining viable and recruitment declines.
To slow or limit the invasion of these exotic cool-season grasses or other exotic plants, it is imperative that managerial techniques (e.g., prescribed grazing, prescribed burning) be carefully constructed, monitored, and evaluated with respect to that objective. If management does not include measures to control or reduce these exotic plants, the transition to State 3: Invaded State should be expected (T2A). Managers need to understand when the plant community is at or near these parameters, and all data available needs to be evaluated to determine needed management actions.
State 3: Invaded State. The threshold for this state is reached when both the exotic cool-season grasses (e.g., Kentucky bluegrass, crested wheatgrass, annual bromes) exceed 30% of the plant community and native grasses represent less than 40% of the community. One community phase has been identified for this state. Managers need to understand when the plant community is at or near these parameters, and all data available needs to be evaluated to determine needed management actions.
The exotic cool-season grasses can be quite invasive and often form monotypic stands. As they increase, both forage quantity and quality of the annual production becomes increasingly restricted to late spring and early summer, even though annual production may increase. Forb diversity often declines. Under non-use or minimal use management, mulch can increase and become a physical barrier to plant growth; this alters nutrient cycling, infiltration, and soil biological activity. As such, desirable native plants become increasingly displaced.
Once the state is well established, prescribed burning and prescribed grazing techniques have been largely ineffective in suppressing or eliminating the exotic cool-season grasses even though some short-term reductions may appear successful. However, assuming there is an adequate component of native grasses to respond to treatments, a restoration pathway to State 2: Native/Invaded State may be accomplished with the implementation of long-term prescribed grazing in conjunction with prescribed burning and possibly a range planting (R3A).
State 4: Invaded Conifer State. This state historically existed as small patches of trees and/or shrubs scattered across the site when precipitation, fire frequency, and other factors enabled woody species to colonize or encroach on the site. This often resulted in a mosaic of patches of woody vegetation interspersed within the grass dominated vegetation.
One community phase has been identified and often results from extended periods of non-use or very light grazing and no fire (T2B, T3A). Prescribed burning and/or chemical/mechanical brush management may lead to State 2: Native/Invaded State or State 3 Invaded State depending upon the amount of exotic cool-season grasses (R4A, R4B).
A marked increase in non-use management and active fire suppression since European influence has enabled this state to expand and become more widespread. This is particularly important to the fire- intolerant juniper species’ ability to expand and exploit and dominate grasslands. Where a conifer seed source is available, woody encroachment processes begin to dominate as fire frequency decreases or fire is eliminated from the site. As depicted in the following diagram, conifer seeds disperse into an intact grassland State 1.0 or 2.0, beginning the process of woody encroachment. Decreased fire frequencies allow conifer to establish allowing for a transition to State 5: Invaded Conifer State.
State 5: Go-Back State may result following cropland abandonment and consists of only one plant community phase. This weedy assemblage may include noxious weeds that need control. Over time, the exotic cool-season grasses (Kentucky bluegrass, crested wheatgrass, and/or annual bromes) will likely predominate.
Initially, due to extensive bare ground and a preponderance of shallow-rooted annual plants, the potential for soil erosion is high. Plant species richness may be high; but overall diversity (i.e., equitability) is typically low, with the site dominated by a relatively small assemblage of species. Due to the lack of native perennials and other factors, restoring the site with the associated ecological processes is difficult. However, a successful range planting may result in something approaching State 2: Native/Invaded State (R5A). Following seeding, prescribed grazing, prescribed burning, haying, and the use of herbicides will generally be necessary to achieve the desired result and control weeds, some of which may be noxious weeds. A failed range planting and/or secondary succession will lead to State 3: Invaded State (R5B).
The following state and transition model diagram illustrates the common states, community phases, community pathways, and transition and restoration pathways that can occur on the site. These are the most common plant community phases and states based on current knowledge and experience; changes may be made as more data are collected. Pathway narratives describing the site’s ecological dynamics reference various management practices (e.g., prescribed grazing, prescribed fire, brush management, herbaceous weed treatment) which, if properly designed and implemented, will positively influence plant community competitive interactions. The design of these management practices will be site specific and should be developed by knowledgeable individuals; based upon management goals and a resource inventory; and supported by an ongoing monitoring protocol.
When the management goal is to maintain an existing plant community phase or restore to another phase within the same state, modification of existing management to ensure native species have the competitive advantage may be required. To restore a previous state, the application of two or more management practices in an ongoing manner will be required. Whether using prescribed grazing, prescribed burning, or a combination of both with or without additional practices (e.g., brush management), the timing and method of application needs to favor the native species over the exotic species. Adjustments to account for variations in annual growing conditions and implementing an ongoing monitoring protocol to track changes and adjust management inputs to ensure desired outcome will be necessary.
The plant community phase composition table(s) has been developed from the best available knowledge including research, historical records, clipping studies, and inventory records. As more data are collected, plant community species composition and production information may be revised.
State and transition model

Figure 10. Rotational landslides are caused by the downward and outward movement of a mass on top of a concave upward failure surface (Abbott 2004).



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Ecosystem states
States 2 and 5 (additional transitions)
T1A | - | Introduction of exotic cool-season grasses |
---|---|---|
T2A | - | Extended periods of non-use or very light grazing, no fire |
T2B | - | Extended periods of non-use or very light grazing, no fire |
R3A | - | Long-term prescribed grazing and prescribed burning with possible range planting |
T3A | - | Extended periods of non-use or very light grazing, no fire |
R4A | - | Prescribed burning and/or chemical/mechanical brush management |
R4B | - | Prescribed burning and/or chemical/mechanical brush management |
R5A | - | Successful range planting with prescribed grazing and prescribed burning |
R5B | - | Failed range planting and/or secondary succession |
T6A | - | Elimination of annual cropping |
State 1 submodel, plant communities
1.1A | - | Soil erosion and deposition |
---|---|---|
1.1B | - | Long-term occupation by prairie dogs |
1.2A | - | Soil stabilization |
1.2B | - | Long-term occupation by prairie dogs |
1.3A | - | Removal of prairie dogs |
State 2 submodel, plant communities
2.1A | - | Soil erosion and deposition |
---|---|---|
2.1B | - | Long-term prairie dog occupation |
2.2A | - | Soil stabilization |
2.3A | - | Prairie dog abandonment/removal |
State 3 submodel, plant communities
State 4 submodel, plant communities
State 5 submodel, plant communities
State 1
Reference State
This state represents the natural range of variability that dominated the dynamics of this ecological site prior to European influence. The primary disturbance mechanisms for this site in the reference condition included soil erosion and deposition (due to landscape position), frequent fire, and grazing by large herding ungulates. Timing of fires and grazing, combined with weather events and erosion/deposition, dictated the dynamics that occurred within the natural range of variability. These factors likely caused the community to shift both spatially and temporally between three community phases. This state was dominated by cool-season grasses with a strong secondary component of short- statured warm-season grasses. Lower fire frequency and intensity permitted fire-intolerant shrubs, such as big sagebrush, to establish and persist. Plant Community Phase 1.1 and 1.2 occurred as a mosaic across this site depending upon the erosional patterns due to runoff from the adjoining, upslope Badland ecological site. Prairie dogs may have occupied and influenced the entire site and shifted both plant community phases to Plant Community Phase 1.3.
Characteristics and indicators. Because of changes in disturbances and other environmental factors (particularly the widespread occurrence of exotic species), the Reference State is considered to no longer exist.
Resilience management. If intact, the reference state should probably be managed with current disturbance regimes which has permitted the site to remain in reference condition, as well as maintaining the quality and integrity of associated ecological sites. Maintenance of the reference condition is contingent upon a monitoring protocol to guide management.
Community 1.1
Western Wheatgrass-Green Needlegrass-Blue Grama (Pascopyrum smithii-Nassella viridula-Bouteloua gracilis): Stabilized
This community phase was historically the most dominant both temporally and spatially. Cool- season grasses dominated the site, but warm-season shortgrasses were also prevalent. Western wheatgrass was the dominant grass. Other grasses and grass-like plants included blue grama, needle and thread, green needlegrass, buffalograss, Sandberg bluegrass, saltgrass, and sedges. Common forbs included scarlet globemallow, upright prairie coneflower, blacksamson echinacea, common sunflower, and blazing star. Common shrubs included silver sagebrush, winterfat, big sagebrush, prairie rose, and prairie sagewort. Annual production varied from about 800-1600 pounds per acre with grasses and grass-like species, forbs, and shrubs, contributing about 85%, 10% and 5%, respectively. Both warm-season grasses and cool-season grasses were well represented in the community. As a result, production was distributed throughout the growing season. This community represents the plant community phase upon which interpretations are primarily based; it is described in the “Plant Community Composition and Group Annual Production” portion of this ecological site description.
Figure 11. Annual production by plant type (representative values) or group (midpoint values)
Table 5. Annual production by plant type
Plant type | Low (lb/acre) |
Representative value (lb/acre) |
High (lb/acre) |
---|---|---|---|
Grass/Grasslike | 710 | 1062 | 1410 |
Forb | 80 | 102 | 125 |
Shrub/Vine | 10 | 36 | 65 |
Total | 800 | 1200 | 1600 |
Community 1.2
Blue Grama-Western Wheatgrass/ Bare Ground (Bouteloua gracilis-Pascopyrum smithii-Bare Ground): Active Erosion/ Deposition
This community phase was the result of soil erosion/deposition of slope alluvium from adjacent Badland ecological sites. Heavy grazing, particularly during droughts, compounded the effects of soil movement. Soil deposition from upslope sites buried existing vegetation which reduced plant vigor and favored more disturbance tolerant species. The slope alluvium was high in calcium carbonate, which is a salt of low solubility but is tolerated by specific grasses and forbs. Needlegrasses and western wheatgrass have declined while blue grama has increased compared to Community Phase 1.1. Associated grasses included Fendler threeawn and other perennial and annual native grasses. Common forbs and shrubs included white sagebrush, false pennyroyal, spiny phlox, woolly plantain, scarlet globemallow, yellow salsify, prairie rose, silver sagebrush, western snowberry, and prairie sagewort. Annual production has decreased compared to Community Phase 1.1.
Community 1.3
Annual Forbs/Bare Ground/ Blue Grama/ Fetid Marigold (Annual Forbs/ Bare Ground/ Bouteloua gracilis/ Dyssodia papposa): Prairie Dog Town
This community phase formed during periods of long-term occupation by prairie dogs. It was characterized by the abundance of annual forbs (e.g., fetid marigold, woolly plantain) and bare ground. Some perennial native species remained but were greatly reduced in vigor and may not have been readily visible.
Pathway 1.1A
Community 1.1 to 1.2
Community Phase Pathway 1.1 to 1.2 resulted from soil erosion/deposition due to its location adjacent to and downslope from the Badland ecological site. This resulted in marked increases in blue grama and bare ground with a corresponding decrease in green needlegrass. The site received additional moisture as well as sediment from the upslope site. Moisture moved across the site as sheet flow or in concentrated flow patterns with minimal benefit to plant growth. Soil erosion would have been evident within concentrated flow patterns while soil deposition may have occurred in areas with sheet flow, depending upon rainfall intensity and frequency.
Pathway 1.1B
Community 1.1 to 1.3
Community Phase Pathway 1.1 to 1.3 resulted from long-term occupation by prairie dogs. This resulted in marked increases in annual forbs, bare ground, blue grama, and fetid marigold along with corresponding decreases in perennial grasses and forbs. Reduced vegetative cover resulting from drought and/or heavy grazing may have facilitated this pathway.
Pathway 1.2A
Community 1.2 to 1.1
Community Phase Pathway 1.2 to 1.1 occurred during periods of soil stabilization. This resulted in marked decreases in blue grama and bare ground with a corresponding increase in green needlegrass. This pathway was initiated by a shift in the waterflow patterns from the adjacent Badland ecological site, possibly due to rotational slides (aka slumping). Flow patterns shifted across this site in response to changing erosional patterns on the Badland Fan ecological site. As these erosional patterns shifted, Plant Community Phases 1.1 and 1.2 may also have shifted locations across the site in response to these new erosion/deposition patterns. Due to the inherit instability of the adjacent upslope Badland landscape, this erosional/depositional pattern should be considered natural. However, long term drought followed by a return to average precipitation, heavy season-long grazing, or a combination of these two disturbances may be contributing factors.
Pathway 1.2B
Community 1.2 to 1.3
Community Phase Pathway 1.2 to 1.3 occurred during periods of long-term occupation by prairie dogs. This resulted in marked increases in annual forbs, bare ground, blue grama, and fetid marigold along with corresponding decreases in perennial grasses and forbs.
Pathway 1.3A
Community 1.3 to 1.2
Community Phase Pathway 1.3 to 1.2 occurred with the abandonment of prairie dogs. This resulted in marked increases in perennial grasses and forbs along with corresponding decreases in annual forbs, bare ground, blue grama, and fetid marigold.
State 2
Native/Invaded State
This state is similar to State 1: Reference State but has now been colonized by the exotic cool-season grasses (commonly Kentucky bluegrass, crested wheatgrass, and/or annual bromes). Although the state is still dominated by native grasses, an increase in these exotic cool-season grasses can be expected. Silver sagebrush is occasionally a conspicuous component of this state. These exotic cool-season grasses can be quite invasive on the site and are particularly well adapted to heavy grazing. They also often form monotypic stands. As these exotic cool-season grasses increase, both forage quantity and quality become increasingly restricted to late spring and early summer due to the monotypic nature of the stand, even though annual production may increase. Native forbs generally decrease in production, abundance, diversity, and richness compared to that of State 1: Reference State. These exotic cool-season grasses have been particularly and consistently invasive under extended periods of no use and no fire. To slow or limit the invasion of these exotic grasses, it is imperative that managerial techniques (e.g., prescribed grazing, prescribed burning) be carefully constructed, monitored, and evaluated with respect to that objective. If management does not include measures to control or reduce these exotic cool-season grasses, the transition to State 3: Invaded State should be expected. Annual production of this state can be quite variable, in large part due to the amount of exotic cool- season grasses. However, as the exotic cool-season grasses increase, peak production will shift to earlier in the growing season.
Characteristics and indicators. The presence of trace amounts of exotic cool-season grasses indicates a transition from State 1 to State 2. The presence of exotic biennial or perennial leguminous forbs (i.e., sweet clover, black medic) may not, on their own, indicate a transition from State 1 to State 2 but may facilitate that transition.
Resilience management. To slow or limit the invasion of these exotic grasses, it is imperative that managerial techniques (e.g., prescribed grazing, prescribed burning) be carefully constructed, monitored, and evaluated with respect to that objective. Grazing management should be applied that enhances the competitive advantage of native grass and forb species. This may include: (1) early spring grazing when exotic cool-season grasses are actively growing and native cool-season grasses are dormant; (2) applying proper deferment periods allowing native grasses to recover and maintain or improve vigor; (3) adjusting overall grazing intensity to reduce excessive plant litter (above that needed for rangeland health indicator #14 – see Rangeland Health Reference Worksheet); (4) incorporating early heavy spring utilization which focuses grazing pressure on exotic cool-season grasses and reduces plant litter, provided that livestock are moved when grazing selection shifts from exotic cool-season grasses to native grasses. Prescribed burning should be applied in a manner that maintains or enhances the competitive advantage of native grass and forb species. Prescribed burns should be applied as needed to adequately reduce/remove excessive plant litter and maintain the competitive advantage for native species. Timing of prescribed burns (spring vs. summer vs. fall) should be adjusted to account for differences in annual growing conditions and applied during windows of opportunity to best shift the competitive advantage to the native species.
Community 2.1
Western Wheatgrass-Green Needlegrass-Blue Grama (Pascopyrum smithii-Nassella viridula-Bouteloua gracilis): Stabilized

Figure 12. Plant Community Phase 2.1, Western Wheatgrass-Green Needlegrass-Blue Grama/Stabilized
This Community Phase is similar to Community Phase 1.1 but has been colonized by exotic cool- season grasses (often Kentucky bluegrass, crested wheatgrass, and/or annual bromes). However, these exotics are present in smaller amounts with the community still dominated by native grasses. Exotic forbs (e.g., Canada thistle, leavy spurge, hound’s tongue) are also known to invade the site.
Community 2.2
Blue Grama-Western Wheatgrass/Bare Ground/ Exotic Cool- Season Grasses (Bouteloua gracilis-Pascopyrum smithii-Bare Ground/Exotic Cool- Season Grasses): Active Erosion/ Deposition

Figure 13. Looking upslope at Community Phase 2.2: Blue Grama-Western Wheatgrass/Bare Ground/Exotic Grasses/Exotic Grasses: Active Erosion/Deposition. Foreground indicates soil deposition from Badland ecological site.

Figure 14. Looking upslope at Community Phase 2.2. Looking upslope, foreground indicates soil deposition and active water flow pattern from Badland ecological site with rotational slide.
This Community Phase is similar to Community Phase 1.2 but has been colonized by exotic cool- season grasses (often Kentucky bluegrass, crested wheatgrass, and/or annual bromes). However, these exotics are present in smaller amounts with the community still dominated by native grasses. Exotic forbs (e.g., Canada thistle, leavy spurge, hound’s tongue) are also known to invade the site. This Community Phase results from soil erosion/deposition due to the landscape position being located downslope from the Badland ecological site. Soil erosion/ deposition are the primary disturbance factors; however, continuous grazing or over-utilization during extended drought periods will compound the effects of soil movement. Soil deposition from upslope sites buries existing vegetation, reducing plant vigor and favoring more disturbance tolerant species. The slope alluvium is high in calcium carbonate, which is a salt of low solubility but is tolerated by specific grasses and forbs. Needlegrasses and western wheatgrass have declined while blue grama has increased compared to Community Phase 1.1. Associated grasses include Fendler threeawn and other perennial and annual native grasses. Common forbs and shrubs include white sagewort, false pennyroyal, spiny phlox, woolly plantain, scarlet globemallow, yellow salsify, prairie rose, silver sagebrush, western snowberry, and prairie sagewort. Annual production has decreased compared to Community Phase 1.1.
Community 2.3
Annual Forbs/ Bare Ground/ Blue Grama/ Fetid Marigold (Annual Forbs/ Bare Ground/ Bouteloua gracilis/ Dyssodia papposa): Prairie Dog Town

This plant community phase is characterized by annual forbs (e.g., fetid marigold, woolly plantain) and bare ground. Some perennial native and exotic species remain but are greatly reduced in vigor and may not be readily visible. Compared to Plant Community Phase 2.1, plant litter and infiltration rate have decreased while bare ground, runoff, and soil erosion have increased.
Pathway 2.1A
Community 2.1 to 2.2


Community Phase Pathway 2.1 to 2.2 results from soil erosion/deposition due to its location adjacent to and downslope from the Badland ecological site. This results in marked increases in blue grama and bare ground with a corresponding decrease in green needlegrass. This site is located adjacent to and downslope from the Badland ecological site. This results in the site receiving additional moisture as well as sediment from the upslope site. Moisture moves across the site as sheet flow or in concentrated flow patterns with minimal benefit to plant growth. Soil erosion will be evident within concentrated flow patterns while soil deposition may occur in areas with sheet flow, depending upon rainfall intensity and frequency.
Pathway 2.1B
Community 2.1 to 2.3


Community Phase Pathway 2.1 to 2.3 results from long-term occupation by prairie dogs. This results in marked increases in annual forbs, bare ground, blue grama, and fetid marigold along with corresponding decreases in perennial grasses and forbs. Reduced vegetative cover resulting from drought and/or heavy grazing may facilitate this pathway.
Pathway 2.2A
Community 2.2 to 2.1


Community Phase Pathway 2.2 to 2.1 occurs during periods of soil stabilization. This results in marked decreases in blue grama and bare ground with a corresponding increase in green needlegrass. This pathway is initiated by a shift in the waterflow patterns from adjacent Badland ecological sites, possibly due to rotational slides (aka slumping). Flow patterns shift across this site in response to changing erosional patterns on the Badland Fan ecological site. As these erosional patterns shift, Plant Community Phases 2.1 and 2.2 may also shift locations across the site in response to these new erosion/deposition patterns. Due to the inherit instability of the adjacent upslope Badland ecological sites, this erosional/depositional pattern would be considered natural. However, long-term drought followed by a return to normal precipitation, heavy season-long grazing, or a combination of these two disturbances may be contributing factors.
Pathway 2.3A
Community 2.3 to 2.2


Community Phase Pathway 2.3 to 2.2 is initiated by prairie dog abandonment or removal and application of prescribed grazing with adequate recovery periods between grazing events.
State 3
Invaded State
This state is the result of invasion and dominance of exotic cool-season grasses. This state is characterized by exotic cool-season grasses (such as Kentucky bluegrass, crested wheatgrass, and/or annual bromes). Once the state is well established, single disturbance events (such as high-intensity fires, severe grazing, or drought) will not result in more than a very short-term reduction of these species.
Characteristics and indicators. This site is characterized by exotic cool-season grasses constituting greater than 30 percent of the annual production and native grasses constituting less than 40 percent of the annual production
Resilience management. Light or moderately stocked continuous, season-long grazing or a prescribed grazing system which incorporates adequate deferment periods between grazing events and proper stocking rate levels will maintain this State. Application of herbaceous weed treatment, occasional prescribed burning and/or brush management may be needed to manage noxious weeds and increasing shrub (e.g., western snowberry) populations.
Community 3.1
Exotic Cool-Season Grasses/ Western Wheatgrass/Forbs (Exotic Cool-Season Grasses/ Pascopyrum smithii/ Forbs)

Figure 15. Plant Community Phase 3.1: Exotic Grasses/Western Wheatgrass/Forbs – Dominated by crested wheatgrass. Note silver sagebrush can occur on this site.
This plant community develops after an extended period (i.e., 10 or more years) of non-use by herbivores and exclusion of fire. Exotic grasses (such as Kentucky bluegrass, crested wheatgrass, and/or annual bromes) tend to invade and may dominate this plant community. Other grasses present in reduced amounts often include western wheatgrass, needle and thread, and Fendler threeawn. Blue grama has declined due to the competition and shading by the exotic cool-season grasses. Common forbs include sweetclover, tarragon, white sagebrush, and American vetch. Prairie sagewort and silver sagebrush may be common shrubs.
State 4
Invaded Conifer State
This state is characterized by a dominance of Rocky Mountain juniper in all layers with scattered mature/decadent green ash trees. Other junipers (e.g., common juniper, creeping juniper, eastern red cedar) and ponderosa pine may also be present. Depending upon juniper density, the herbaceous layer may be minimal to non-existent and dominated by needlecast. The hydrologic function of this state has changed compared to that of State 1: Reference State and State 2: Native/Invaded State. The shallow rooting structure of the juniper and lack of native perennial grass species has reduced infiltration and increased the potential of erosion.
Characteristics and indicators. The dominance of woody species (by cover and production) distinguishes this state from other herbaceously dominated states.
Resilience management. This state is resistant to change in the long-term absence of fire. Restoration efforts would require the use of prescribed fire, mechanical treatment, and prescribed grazing. Considerable time and effort will be required to restore to other States.
Community 4.1
Juniper/ Pine Species (Juniperus spp./ Pinus ponderosa)

Figure 16. Foreground – Plant Community Phase 4.1 Juniper/Pine – dominated by Rocky Mountain Juniper
This community is characterized by the complete dominance of junipers in all layers of the community (i.e., tree, sapling, shrub, and herbaceous). Ponderosa pine may also be a conspicuous component of the community. Rocky Mountain juniper is generally the most abundant juniper, but common juniper, creeping juniper, and eastern red cedar may also be present. Green ash is often present but in low densities and only in a mature state. Dependent upon juniper density, chokecherry, American plum, snowberry, Saskatoon serviceberry, and other shrubs are often sparse or absent. Herbaceous vegetation is generally sparse to absent. A marked increase in non-use management and active fire suppression since European influence has enabled this state to expand and become more widespread.
State 5
Go-Back State
This state is highly variable depending on the level and duration of disturbance related to the T6A transitional pathway. In this MLRA, the most probable origin of this state is plant succession following cropland abandonment or, perhaps, recreational activity or concentrated livestock activity for a prolonged period. This plant community will initially include a variety of annual forbs and grasses, some of which may be noxious weeds and need control. Over time, the exotic cool-season grasses (Kentucky bluegrass, crested wheatgrass, and/or annual bromes) will likely predominate.
Characteristics and indicators. Tillage has destroyed the native plant community, altered soil structure and biology, reduced soil organic matter, and resulted in the formation of a tillage induced compacted layer which is restrictive to root growth. Removal of perennial grasses and forbs results in decreased infiltration and increased runoff.
Resilience management. Continued tillage will maintain the state. Control of noxious weeds will be required.
Community 5.1
Annual/Pioneer-Perennial/Exotic Species
Tillage has destroyed the native plant community, altered soil structure and biology, reduced soil organic matter, and resulted in the formation of a tillage induced compacted layer which is restrictive to root growth. Removal of perennial grasses and forbs results in decreased infiltration and increased runoff.
State 6
Any Plant Community
Transition T1A
State 1 to 2
This is the transition from the State 1: Reference State to the State 2: Native/Invaded State due to the introduction and establishment of exotic cool-season grasses (typically Kentucky bluegrass, crested wheatgrass, and/or annual bromes). This transition was probably inevitable and corresponded to a decline in native warm-season and cool-season grasses; it may have been exacerbated by chronic season-long or heavy late season grazing. Complete rest from grazing and suppression of fire could also have hastened the transition. The threshold between states was crossed when Kentucky bluegrass, crested wheatgrass, annual bromes, or other exotic species became established on the site.
Constraints to recovery. Current knowledge and technology will not facilitate a successful restoration to Reference State.
Transition T2A
State 2 to 3
This is the transition from State 2: Native/Invaded State to State 3: Invaded state resulting from extended periods of non-use or very light grazing, and no fire. Complete rest or light grazing (less than 20 percent utilization) and elimination of fire are the two major contributors to this transition. Exotic cool-season grasses (commonly Kentucky bluegrass, crested wheatgrass, and/or annual bromes) become the dominant graminoids. Preliminary studies indicate this threshold may exist when exotic cool-season grasses exceed 30 percent of the plant community while native grasses represent less than 40 percent of the plant community composition.
Constraints to recovery. Variations in growing conditions (e.g., cool, wet spring) will influence effects of various management activities on exotic cool-season grass populations.
Transition T2B
State 2 to 4
This transition from the State 2: Native/Invaded to State 4: Invaded Conifer State generally occurs during extended periods of non-use or very light grazing, and no fire. It also frequently occurs when the site is in close proximity to stands of juniper and/or ponderosa pine that can serve as a seed source for these species to colonize the site. It has also become more frequent following European settlement when the historic fire regime was markedly reduced.
Constraints to recovery. The extended fire interval may make recovery doubtful due to the abundance of exotic cool-season grasses and lack of native grasses. Fire intensity along with consumption of available fuels may cause incomplete or patchy burns. Ladder fuel and/or fuel loading are required for successfully controlling ponderosa pine (crown vs. ground fire). Continued recruitment of seeds (juniper and pine) from adjacent sites will hamper site restoration. Constraints to recovery include reticence to undertake tree removal and the perception that trees may be a desirable vegetation component for wildlife habitat, carbon sequestration, aesthetics, etc. Managing the site for mule deer, big horn sheep, livestock, or grassland nesting birds will need to consider the intensive management required to restore and maintain the site in State 2. The disturbance regime necessary to restore this site to State 2: Native/Invaded State is very labor intensive and costly; therefore, addressing woody removal earlier in the encroachment phase is the most cost-effective treatment for woody control.
Restoration pathway R3A
State 3 to 2
This restoration pathway from State 3: Invaded State to State 2: Native/Invaded State may be accomplished with the implementation of long-term prescribed grazing and prescribed burning, assuming there is an adequate component of native grasses to respond to the treatments. Depending upon the abundance of the exotic cool-season grasses, a follow-up range planting may be necessary to complete the restoration. Both prescribed grazing and prescribed burning are likely necessary to successfully initiate this restoration pathway, the success of which depends upon the presence of a remnant population of native grasses in Community Phase 3.1. That remnant population, however, may not be readily apparent without close inspection. The application of several prescribed burns may be needed at relatively short intervals in the early phases of this restoration process, in part because many of the shrubs (e.g., western snowberry) sprout profusely following one burn. Some previous efforts have shown promise with early season prescribed burning; however, summer or fall burning may also be effective under certain circumstances depending on exotic grass species. The prescribed grazing should include adequate recovery periods following each grazing event and stocking levels which match the available resources. If properly implemented, this will shift the competitive advantage from the exotic cool-season grasses to the native cool-season grasses.
Context dependence. Grazing management should be applied in a manner that enhances/maximizes the competitive advantage of native grass and forb species over the exotic species. This may include the use of prescribed grazing to reduce excessive plant litter accumulations above that needed for rangeland health indicator #14 (see Rangeland Health Reference Worksheet). Increasing livestock densities may facilitate the reduction in plant litter provided length and timing of grazing periods are adjusted to favor native species. Grazing prescriptions designed to address exotic grass invasion and favor native species may involve earlier, short, intense grazing periods with proper deferment to improve native species health and vigor. Fall (e.g., September, October) prescribed burning followed by an intensive, early spring graze period with adequate deferment for native grass recovery may shift the competitive advantage to the native species, facilitating the restoration to State 2: Native/Invaded. Prescribed burning should be applied in a manner that enhances the competitive advantage of native grass and forb species over the exotic species. Prescribed burns should be applied at a frequency which mimics the natural disturbance regime, or more frequently as is ecologically (e.g., available fuel load) and economically feasible. Burn prescriptions may need adjustment to: (1) account for change in fine fuel orientation (e.g., “flopped” Kentucky bluegrass); (2) fire intensity and duration by adjusting ignition pattern (e.g., backing fires vs head fires); (3) account for plant phenological stages to maximize stress on exotic species while favoring native species (both cool- and warm-season grasses).
Transition T3A
State 3 to 4
This transition from the State 3: Invaded Grass State to State 4: Invaded Conifer State generally occurs during extended periods of non-use or very light grazing, and no fire. It frequently occurs when the site is in close proximity to junipers and/or ponderosa pines where they can serve as a seed source for these species to colonize the site. It has also become more frequent following European settlement when the historic fire regime was markedly reduced.
Constraints to recovery. Lack of fine fuels in the tree understory limits fire intensity causing incomplete or patchy burns. Ladder fuels and/or fuel loading are required, but are limited, for successfully controlling ponderosa pine (crown vs. ground fire). Cost of mechanical and/or chemical treatment may be prohibitive. Continued recruitment of seeds (juniper and pine) from adjacent sites and sprouting nature of deciduous trees and shrubs will hamper site restoration. Existing herbaceous vegetation (native grasses and cool-season exotics) may be lacking allowing for invasive species (e.g., Canada thistle, leafy spurge) to dominate after a prescribed fire. Constraints to recovery include reticence to undertake tree removal and the perception that trees may be a desirable vegetation component for wildlife habitat, carbon sequestration, aesthetics, etc. Managing the site for mule deer, big horn sheep, livestock, or grassland nesting birds will need to consider the intensive management required to restore and maintain the site in State 2. The disturbance regime necessary to restore this site to State 2: Native/Invaded State is very labor intensive and costly; therefore, addressing woody removal earlier in the encroachment phase is the most cost-effective treatment for woody control.
Restoration pathway R4A
State 4 to 2
This restoration from State 4: Invaded Conifer State to State 2: Native/Invaded State results from prescribed burning and/or chemical/mechanical brush management. This assumes that the exotic cool-season grasses do not dominate the understory. If exotic cool-season grasses dominate the restoration effort will lead to State 3: Invaded State (via R4B).
Context dependence. Fire intensity along with consumption of available fuels may cause incomplete or patchy burns. Ladder fuel and/or fuel loading are required for successfully controlling ponderosa pine (crown vs. ground fire). Continued recruitment of seeds (juniper and pine) from adjacent sites will hamper site restoration. Intensive management is required to restore and maintain the site in State 2: Native/Invaded State.
Restoration pathway R4B
State 4 to 3
This restoration from State 4: Invaded Conifer State to State 3: Invaded State results from prescribed burning and/or chemical/mechanical brush management. This assumes that exotic cool-season grasses dominate the understory. If native grasses dominate the understory the restoration effort will lead to State 2: Native/Invaded State (via R4A).
Context dependence. Fire intensity along with consumption of available fuels may cause incomplete or patchy burns. Ladder fuel and/or fuel loading are required for successfully controlling ponderosa pine (crown vs. ground fire). Continued recruitment of seeds (juniper and pine) from adjacent sites will hamper site restoration. Intensive management is required to restore and maintain the site in State 2: Native/Invaded State.
Restoration pathway R5A
State 5 to 2
This Restoration Pathway from State 5: Go-Back State to State 2: Native/Invaded State can be accomplished with a successful range planting. Following seeding, prescribed grazing, prescribed burning, haying, or use of herbicides will generally be necessary to achieve the desired result and control any noxious weeds. It may be possible using selected plant materials and agronomic practices to approach something very near the functioning of State 2: Native/Invaded State. Application of chemical herbicides and the use of mechanical seeding methods using adapted varieties of the dominant native grasses are possible and can be successful. After establishment of the native plant species, prescribed grazing should include adequate recovery periods following each grazing event and stocking levels which match the available resources; management objectives must include the maintenance of those species, the associated reference state functions, and continued treatment of exotic grasses.
Context dependence. A successful range planting will include proper seedbed preparation, weed control (both prior to and after the planting), selection of adapted native species representing functional/structural groups inherent to the State 1, and proper seeding technique. Management (e.g., prescribed grazing, prescribed burning) during and after establishment must be applied in a manner that maintains the competitive advantage for the seeded native species. Adding non-native species can impact the above and below ground biota. Elevated soil nitrogen levels have been shown to benefit smooth brome and Kentucky bluegrass more than some native grasses. As a result, fertilization, exotic legumes in the seeding mix, and other techniques that increase soil nitrogen may promote smooth brome and Kentucky bluegrass invasion. The method or methods of herbaceous weed treatment will be site specific to each situation; but generally, the goal would be to apply the pesticide, mechanical control, or biological control (either singularly or in combination) in a manner that shifts the competitive advantage from the targeted species to the native grasses and forbs. The control method(s) should be as specific to the targeted species as possible to minimize impacts to non-target species.
Restoration pathway R5B
State 5 to 3
This Restoration Pathway from State 5: Go-Back State to State 3: Invaded State results from a failed range planting and/or secondary succession.
Context dependence. Failed range plantings can result from many causes (both singularly and in combination) including drought, poor seedbed preparation, improper seeding methods, seeded species not adapted to the site, insufficient weed control, herbicide carryover, poor seed quality (purity & germination), and/or improper management.
Restoration pathway T6A
State 6 to 5
This transition is from any plant community to State 5: Go-Back State. It is most commonly associated with the cessation of cropping without the benefit of range planting, resulting in a “go-back” situation. Soil conditions can be quite variable on the site, in part due to variations in the management/cropping history (e.g., development of a tillage induced compacted layer (plow pan), erosion, fertility, and/or herbicide/pesticide carryover). Thus, soil conditions should be assessed when considering restoration techniques.
Additional community tables
Table 6. Community 1.1 plant community composition
Group | Common name | Symbol | Scientific name | Annual production (lb/acre) | Foliar cover (%) | |
---|---|---|---|---|---|---|
Grass/Grasslike
|
||||||
1 | Cool-season grasses | 420–540 | ||||
western wheatgrass | PASM | Pascopyrum smithii | 240–360 | – | ||
green needlegrass | NAVI4 | Nassella viridula | 120–240 | – | ||
needle and thread | HECOC8 | Hesperostipa comata ssp. comata | 60–120 | – | ||
2 | Warm-season grasses | 180–300 | ||||
blue grama | BOGR2 | Bouteloua gracilis | 180–300 | – | ||
buffalograss | BODA2 | Bouteloua dactyloides | 12–60 | – | ||
3 | Other native grasses | 60–180 | ||||
Grass, perennial | 2GP | Grass, perennial | 12–60 | – | ||
Sandberg bluegrass | POSE | Poa secunda | 12–60 | – | ||
prairie Junegrass | KOMA | Koeleria macrantha | 12–60 | – | ||
plains muhly | MUCU3 | Muhlenbergia cuspidata | 12–60 | – | ||
saltgrass | DISP | Distichlis spicata | 12–60 | – | ||
Fendler threeawn | ARPUL | Aristida purpurea var. longiseta | 12–36 | – | ||
sand dropseed | SPCR | Sporobolus cryptandrus | 0–24 | – | ||
Grass, annual | 2GA | Grass, annual | 0–12 | – | ||
4 | Grass-Likes | 24–60 | ||||
threadleaf sedge | CAFI | Carex filifolia | 12–60 | – | ||
Grass-like (not a true grass) | 2GL | Grass-like (not a true grass) | 0–36 | – | ||
needleleaf sedge | CADU6 | Carex duriuscula | 12–24 | – | ||
Forb
|
||||||
5 | Forbs | 84–120 | ||||
blacksamson echinacea | ECAN2 | Echinacea angustifolia | 12–24 | – | ||
common sunflower | HEAN3 | Helianthus annuus | 12–24 | – | ||
stiffstem flax | LIRI | Linum rigidum | 12–24 | – | ||
Indian breadroot | PEDIO2 | Pediomelum | 12–24 | – | ||
upright prairie coneflower | RACO3 | Ratibida columnifera | 12–24 | – | ||
Forb (herbaceous, not grass nor grass-like) | 2FORB | Forb (herbaceous, not grass nor grass-like) | 0–24 | – | ||
Forb, perennial | 2FP | Forb, perennial | 0–24 | – | ||
scarlet globemallow | SPCO | Sphaeralcea coccinea | 12 | – | ||
white sagebrush | ARLU | Artemisia ludoviciana | 0–12 | – | ||
rough false pennyroyal | HEHI | Hedeoma hispida | 0–12 | – | ||
spiny phlox | PHHO | Phlox hoodii | 0–12 | – | ||
woolly plantain | PLPA2 | Plantago patagonica | 0–12 | – | ||
yellow salsify | TRDU | Tragopogon dubius | 0–12 | – | ||
Shrub/Vine
|
||||||
6 | Shrubs | 12–60 | ||||
silver sagebrush | ARCA13 | Artemisia cana | 12–24 | – | ||
prairie sagewort | ARFR4 | Artemisia frigida | 12–24 | – | ||
prairie rose | ROAR3 | Rosa arkansana | 12–24 | – | ||
western snowberry | SYOC | Symphoricarpos occidentalis | 12–24 | – | ||
Shrub (>.5m) | 2SHRUB | Shrub (>.5m) | 0–24 | – | ||
big sagebrush | ARTR2 | Artemisia tridentata | 0–12 | – | ||
winterfat | KRLA2 | Krascheninnikovia lanata | 0–12 | – |
Interpretations
Animal community
Animal Community – Wildlife Interpretations
Landscape
The MLRA 58C landscape is characterized by moderately dissected rolling plains with areas of local Badlands, buttes, and isolated hills. MLRA 58C is considered to have a continental climate with cold winters and hot summers, low humidity, light rainfall, and much sunshine. Extremes in temperature are common and characteristic of the MLRA. This area supports natural mixed-grass prairie vegetation with prairie rose, leadplant, and patches of western snowberry interspersed throughout the area. Green ash, chokecherry, and buffaloberry occur in draws and narrow valleys, creating woody riparian corridors. Complex/intermingled ecological sites create diverse grass- and shrubland habitats interspersed with varying densities linear, slope, depressional, and in-stream wetlands associated with headwater streams and tributaries to the Missouri River. These habitats provide critical life-cycle components for many wildlife species.
Historic Communities/Conditions within MLRA 58C:
The northern mixed-grass prairie was a disturbance-driven ecosystem with fire, herbivory, and climate functioning as the primary ecological drivers either singly or often in combination. Many species of grassland birds, small mammals, insects, reptiles, amphibians, and large herds of roaming bison, elk, and pronghorn were historically among the inhabitants adapted to this semi-arid region. Roaming herbivores, as well as several small mammal and insect species, were the primary consumers linking the grassland resources to large predators (such as the wolf, mountain lion, and grizzly bear) and smaller carnivores (such as the coyote, bobcat, red fox, and raptors). The black-tailed prairie dog was once abundant and provided ecological services by manipulating the plant and soil community providing habitat for the black-footed ferret, burrowing owl, ferruginous hawk, mountain plover, swift fox, small mammals, and amphibians and reptiles. Extirpated species include free-ranging American bison, Canada lynx, common raven, grizzly bear, gray wolf, black-footed ferret, mountain plover, and peregrine falcon (breeding). Extinct from the region is the Rocky Mountain locust.
Present Communities/Conditions within MLRA 58C:
Following European influence, domestic livestock grazing, elimination of fire, energy development, and other anthropogenic factors influenced plant community composition and abundance. Transportation corridors, energy development, and Rocky Mountain juniper and ponderosa pine encroachment are the main factors contributing to habitat fragmentation, reducing habitat quality for area-sensitive species. These influences fragmented the landscape, reduced or eliminated ecological drivers (fire), and introduced exotic plant species including, smooth brome, crested wheatgrass, Kentucky bluegrass, and leafy spurge further impacting plant and animal communities. The loss of the bison, black-tailed prairie dogs, and fire as primary ecological drivers greatly influenced the character of the remaining native plant communities and the associated wildlife moving towards a less diverse and more homogeneous landscape, lacking diverse species composition and stature.
Extensive fragmentation by annual cropping has not occurred within the MLRA. Limited fragmentation from annual cropping or tame hay production has occurred within the Little Missouri River flood plain and the higher, flat plateaus. Fragmentation east and west of MLRA 58C has funneled many species into this area in search of expansive grasslands.
Some wildlife species in this area are: mule deer, white-tailed deer, elk, bighorn sheep, pronghorn, mountain lion, coyote, red fox, bobcat, prairie rattlesnake, American badger, raccoon, North American porcupine, beaver, striped skunk, American mink, white-tailed jackrabbit, black-tailed prairie dog, Eastern and Merriam’s wild turkey, golden eagle, ferruginous hawks, sharp-tailed grouse, greater sage-grouse, black-billed magpie, and numerous species of grassland-nesting birds and pollinating insects.
Presence of wildlife species is often determined by ecological site characteristics including grass and forb species, hydrology, aspect, and other associated ecological sites. Home ranges for most species are larger than one ecological site or are dependent on more than one ecological site for annual life requisites. Ecological sites offer different habitat elements as the annual life requisites change. Habitat improvement and creation must be conducted within the mobility limits of a known population for the species.
Insects play an important role providing ecological services for plant community development. Insects that are scavengers or aid in decomposition provide the food chain baseline sustaining the carnivorous insects feeding upon them. Many insects provide the ecological services necessary for pollination, keeping plant communities healthy and productive. Insects provide a protein food source for numerous species including grassland nesting birds and their young.
Species unique to the MLRA:
Mountain Lions: Mountain lions were absent for many years in MLRA 58C but have recently taken up permanent residency within the region. Mountain lions occur in of the Little Missouri Badlands and woody habitat in MLRA 58C. Rugged terrain and forest provide excellent stalking cover to hunt large mammals and other prey. Mountain lions make use of caves for escape and loafing cover.
Bighorn sheep: Bighorn sheep make use of the rugged terrain, rocky outcrops, and high plateaus of MLRA 58C along the Little Missouri River. North Dakota bighorn sheep populations are almost exclusively within MLRA 58C. Bighorn sheep were once extirpated from North Dakota but were successfully reintroduced in the mid-twentieth century. They now occur in several distinct populations within MLRA 58C. Rocky Mountain juniper encroachment degrades the limited habitat for bighorn sheep. Managers should consider bighorn sheep association with domestic sheep, since transfer of pneumonia and other diseases is known to occur.
Golden eagle: The Badlands within MLRA 58C are key areas for Golden eagle nesting. Adjacent grasslands, shrublands, and black-tailed prairie dog towns are used for hunting.
Bats: Except for hibernacula, MLRA 58C provides life requisites for several bat species, in part due to presence of riparian forest, wooded draws, caves, and rocky outcrops. No known hibernacula have been found in North Dakota.
Short-horned lizard and sagebrush lizard: This MLRA provides preferred habitat for these two species. The short-horned lizard prefers semi-arid, shortgrass prairie in rough terrain; it is uncommon to locally abundant in MLRA 58C. The rare sagebrush lizard prefers sagebrush and rocky areas provided by this MLRA and adjacent MLRA 58D.
Greater sage-grouse and Brewer’s sparrow: The extreme southwest extension of MLRA 58C have ecological sites capable of producing sufficient big sage canopy cover to provide greater sage-grouse life requisites. MLRA 58C and 58D are the only MLRAs in North Dakota that support Wyoming big sage brush (big sage) production. Research data indicates greater sage-grouse prefer big sagebrush canopy cover for nesting at ≥8% with an average height of around 16 inches. The species prefers winter cover canopy that averages 15% with an average height of around 8 inches. Soil site potential, management, climate, and other factors all play a role in the amount, if any, of big sagebrush on an ecological site. Changes in big sage canopy cover occur slowly (30-50 years) unless the site is impacted by fire or cultivation. Big sage recovery after a burn can take 30 to 100 years. Greater sage-grouse and Brewer’s sparrow habitat and populations are reduced or eliminated when big sagebrush canopy is reduced to less than 8% for greater sage-grouse and 10% cover for Brewer’s sparrow. As conifer encroachment increases, greater sage-grouse lekking activity decreases. Once conifer encroachment exceeds 4% canopy cover, no leks remain.
Species of Concern within the MLRA:
Following is a list of species considered “species of conservation priority” in the North Dakota State Wildlife Action Plan (2015); “species of greatest conservation need” in the Montana State Wildlife Action Plan (2015); and species listed as “threatened, endangered, or petitioned” under the Endangered Species Act within MLRA 58C at the time this section was developed:
Invertebrates: Dakota skipper, monarch butterfly, regal fritillary, yellow-banded bumble bee, and western bumble bee
Birds: American kestrel, Baird’s sparrow, bald eagle, black-billed cuckoo, bobolink, Brewer’s sparrow, burrowing owl, chestnut-collared longspur, common poorwill, eastern screech-owl ferruginous hawk, golden eagle, grasshopper sparrow, greater sage-grouse, lark bunting, loggerhead shrike, long-billed curlew, marbled godwit, McCown’s longspur, mountain plover, northern harrier, northern pintail, peregrine falcon (migration), prairie falcon, red knot (migration), red-headed woodpecker, sharp-tailed grouse, short-eared owl, Sprague’s pipit, Swainson’s hawk, upland sandpiper, western meadowlark, Wilson’s phalarope, whooping crane (migration), and willet.
Mammals: Big brown bat, black-footed ferret, black-tailed prairie dog, dwarf shrew, gray wolf, hispid pocket mouse, little brown bat, long-eared bat, long-legged bat, meadow jumping mouse, Merriam’s shrew, northern long-eared bat, porcupine, sagebrush vole, swift fox, Townsend’s big-eared bat, and western small-footed bat.
Amphibians and Reptiles: Common snapping turtle, Great Plains toad, greater short- horned lizard, milk snake, northern leopard frog, plains hognose snake, plains spadefoot, sagebrush lizard, smooth softshell, smooth green snake, and spiny softshell.
Fish and Mussels: Blue sucker, burbot, Flathead chub, northern redbelly dace, sickle-fin chub, pearl dace, shortnose gar, sturgeon chub, and sauger.
Grassland Management for Wildlife in the MLRA:
Management activities within the community phase pathways impact wildlife. Community phase, transitional, and restoration pathways are keys to long-term management within each State and between States. Significant inputs must occur to cross the threshold between States (e.g., State 3.0 to 2.0) requiring substantial economic inputs and management (mechanical, reseeding, prescribed fire, woody vegetation removal, grazing intensity, etc.). Timing, intensity, and frequency of these inputs can have dramatic positive or negative effects on local wildlife species. Ranchers and other land managers must always consider the long-term beneficial effects of management on the habitat in comparison to potential short-term negative effects to individuals.
Ecological sites occur as intermingled complexes on the landscape with gradual or sometimes abrupt transitions. Rarely do ecological sites exist in large enough acreage to manage independently. Ecological sites, supporting a dominance of herbaceous vegetation (Loamy/Limy Residual), can be located adjacent to ecological sites that support medium to tall shrubs (Loamy Overflow). Conversely, ecological sites that are dominated by short to mid statured grasses (Claypan) can be adjacent to sites with bare soil only supporting minor amounts of short grasses and forbs (Thin Claypan).
Management of these complex ecological sites can provide a heterogeneous or a homogenous landscape. Grassland bird use declines as the plant community transitions to a homogenous state. Managers need to recognize ecological sites and the complexes they occur in to properly manage the landscape. A management regime for one ecological site may negatively impact an adjacent site (e.g., alteration of a grazing regime within a Flat Bottom Wooded Draw ecological site to encourage understory growth may encourage exotic, cool-season grasses to increase or dominate an adjacent ecological site).
Life requisites and habitat deficiencies are determined for targeted species. Deficiencies need to be addressed along community phase, transitional, and restoration pathways as presented in specific state and transition models. Ecological sites should be managed and restored within the capabilities of the site to provide sustainable habitat. Managers also need to consider habitat provided by adjacent/intermingled ecological sites for species with home ranges or life requisites that cannot be provided by one ecological site.
With populations of many grassland-nesting birds in decline, it is important to maintain these ecological sites in a 1.0 Reference State or the 2.0 Native/Invaded. Plant communities, optimal for a guild of grassland species, serve as a population source where the birth rate exceeds mortality. Species may use marginal plant communities; however, these sites may function as a population sink where mortality exceeds the birth rate.
Understanding preferred vegetative stature and sensitivity to woody encroachment is necessary to manage for the specific grassland species. Various grass heights may be used for breeding, nesting, foraging, or winter habitat. While most species use varying heights, many have a preferred vegetative stature height. Please reference the provisional ESD on the North Dakota eFOTG (linked below) for a chart that provides preferred vegetative stature heights and sensitivity to woody vegetation encroachment.
https://efotg.sc.egov.usda.gov/references/public/ND/58C_Badland_Fan_Narrative_Final_Ref_FSG.pdf
Badland Fan Wildlife Habitat Interpretation:
Badland Fan ecological sites are affected by deposition of colluvium from adjacent Badland (very steep, barren land consisting of soft sedimentary rocks) from sheet flow or in concentrated flow patterns. Limitations within Badland Fan sites support shorter stature and lower diversity of grasses and forbs for wildlife. Associated ecological sites include Badland, Shallow Loamy, Limy Residual, Thin Claypan, and Very Shallow. This complex of ecological sites provides habitat for many edge-sensitive grassland bird species and prairie dog towns.
Badland Fan habitat features and components commonly support grassland nesting birds, notably sharp-tailed grouse leks. Insects rely on associated forbs and grasses for survival and serve as food sources for birds and their young, and as forage for small and large herbivores.
Badland Fan ecological sites may be found in five plant community states (1.0 Reference State, 2.0 Native/Invaded State, 3.0 Invaded State, 4.0 Invaded Conifer State, and 5.0 Go- Back State) within a local landscape. Multiple plant community phases exist within each state. Today, these states occur primarily in response to grazing and drought. Secondary influences include anthropogenic disturbances, black-tailed prairie dogs, and fire.
Because there is no known restoration pathway from State 2.0 to State 1.0, it is important to intensively manage (using tools in the community phase pathways in States 1.0 and 2.0) to prevent further plant community degradation along either the T1A Transitional Pathway to the Native/Invaded State 2.0 or the T2A Transitional Pathway to the Invaded State 3.0 thresholds. Native wildlife generally benefits from grasslands that are heterogeneous in species composition and stature found in States 1.0 and 2.0 that include diverse grass and forb species with varying stature and density. As plant communities degrade within State 2.0, warm-season grasses (particularly short-statured grasses) increase while native forbs are reduced. This transition results in reduced stature and increased plant community homogeneity. When adjacent and/or intermingled ecological sites undergo the same transition, the result can be an expansive, homogenous landscape.
Success along restoration pathway R3A from State 3.0 to State 2.0 is very difficult and is dependent upon the presence of a remnant native grass population. This concept also applies to wildlife, as the target species must either be present on adjacent State 1.0 or State 2.0 plant communities or on ecological sites within the species’ mobility limits. Species with limited mobility, such as some butterflies, must exist near the plant community to utilize restored sites. Mobile species, such as grassland-nesting birds, can easily locate isolated, restored plant communities.
Plant Community Phase 3.1 shows dramatically increased homogeneity of exotic cool- season grasses and further reduction in native forbs. Reduced forb diversity limits insect populations, negatively affecting grassland-nesting bird foraging opportunities. Increased exotic grass litter can limit access to bare ground to nesting insects and can limit mobility by small chicks. A homogenous grassland landscape does not provide quality escape or winter cover. As a result, many species are not able to meet life requisites.
Management along community phase, transition, or restoration pathways should focus upon attainable changes. Short- and long-term monetary costs must be evaluated against short- and long-term ecological services in creating and maintaining habitat of sufficient quality to support a sustainable population density.
1.0 Reference State
Community Phase 1.1 Western Wheatgrass-Green Needlegrass-Blue Grama:
Stabilized: This plant community offers good wildlife habitat; every effort should be made to maintain this ecological site within this community phase. This phase retains high functionality through continued maintenance including prescribed grazing with adequate recovery period as well as prescribed burning. Predominance of grass species in this community favors grazers and mixed- feeders (animals selecting grasses as well as forbs and shrubs). The structural diversity provides habitat for a wide array of migratory and resident birds.
Invertebrates: Insects play a role in maintaining the forb community and provide a forage base for grassland birds, reptiles, and rodents. Ecological services, historically provided by bison, are mirrored by domestic livestock. These services include putting plant material and dung in contact with mineral soil to be used by lower trophic level consumers (such as invertebrate shredders, predators, herbivores, dung beetles, and fungal-feeders).
Dakota skippers do not prefer this site due to limited host plants, such as little bluestem and prairie dropseed. Nuttall's and prairie violets are uncommon, limiting Regal fritillary habitat. Monarch butterflies may use flowering forbs on this site; however, few milkweed species are found on this site to support caterpillar food. Bumblebees and other native bees utilize season- long forbs as a nectar source and bare ground for nesting amongst bunchgrasses. Prescribed grazing with adequate recovery periods (as well as prescribed fire) to maintain the 1.1 Phase has little effect on nests of ground-dwelling insects.
Birds: This plant community provides quality nesting, foraging, and escape habitats favored by short- to midgrass nesting birds. Plant stature may be too dense or tall for burrowing owl and McCown’s longspur; however, it may be used during periods of drought or management (such as rotational grazing or fire) that results in defoliation along Community Phase Pathway 1.1A.
Several species of grassland birds that prefer short- to mid-grass stature will use this site. In years with reduced precipitation or drought, nesting recruitment may be compromised. This plant community provides suitable areas for sharp-tailed grouse leks, nesting, and brood-rearing habitat with limited winter habitat. Limited stature and diverse prey populations provide good hunting opportunities for grassland raptors.
Mammals: The diversity of grasses and forbs provide high nutrition levels for small and large herbivores including voles, mice, rodents, jackrabbits, pronghorn, and deer (white-tailed and mule). Short- to moderate-statured vegetation provides suitable food and thermal, protective, and escape cover for small herbivores such as the hispid pocket mouse.
Amphibians and Reptiles: This ecological site and associated plant communities provide habitat for smooth green snakes. This ecological site can provide habitat for the northern leopard frog and Great Plains toad if freshwater habitats (such as stock water ponds) are located in or adjacent to the site. Silver sagebrush provides habitat to support sagebrush lizard; however, due to the lack of sandy soils and/or rocky areas, neither the sagebrush lizard nor short-horned lizard may use this ecological site. This ecological site provides limited habitat for the plains hog-nosed snake (prefer sandy soils) and plains spadefoot (prefer gravelly or sandy soils).
Fish and Mussels: This ecological site is not directly associated with streams, rivers, or water bodies. Associated ecological sites, such as Clayey or Limy Residual, can receive run-on hydrology from Badland Fan sites. Management on these interconnected sites will have limited, secondary effects on aquatic species.
Community Phase 1.2 Blue Grama-Western Wheatgrass/Bare Ground: Active Erosion/Deposition: Blue grama and western wheatgrass will dominate due to naturally occurring active soil erosion/deposition. This plant community is 85% grasses and grass-like species, with 10% forbs and 5% shrubs. The dominant forbs include white sagebrush, pennyroyal, spiny phlox, woolly plantain, scarlet globemallow, and yellow salsify. The dominant shrubs are prairie rose, silver sagebrush, western snowberry, and prairie sagewort. However, Wyoming big sage could be present due to the abundance of bare ground and lack of fine fuel to carry a fire. If present, Wyoming big sage will be located along the transition zone between the Badland landform, not providing sufficient habitat to meet greater sage grouse life requisites.
Invertebrates: The combination of forbs and flowering shrubs provides season long pollen and nectar sources for native pollinators. However, this site does not provide life requisites for butterfly species of concern. Abundant bare ground provides nesting sites for ground nesting pollinator species.
Birds: This plant community provides nesting, foraging, and escape habitats favored by short- to mid-grass nesting birds. A shift to shorter herbaceous plant statures and a short shrub component along Community Phase Pathway 1.1A begins to benefit McCown’s longspur, chestnut-collared longspur, horned lark, and burrowing owl. Species that prefer short- to mid- grass statures will be generally successful with normal to above normal precipitation. In years with reduced precipitation or heavy grazing, nesting recruitment may be compromised for midgrass nesting species. This plant community provides areas suitable for sharp-tailed grouse leks. Limited cover and diverse prey populations provide good hunting opportunities for grassland raptors.
Mammals: Provides similar life requisites as Community Phase 1.1.
Amphibians and Reptiles: Provides similar life requisites as Community Phase 1.1.
Fish and Mussels: Provides similar life requisites as Community Phase 1.1.
Community Phase 1.3 Annual Forbs/Bare Ground/Blue Grama/Fetid Marigold (Prairie Dog Town): This plant community phase is characterized by grazing-tolerant species and annual forbs (e.g., fetid marigold). Continued heavy grazing, repeated drought, short-term prairie dog occupation, or a combination of these disturbances will shift to increased annual forbs with a reduction in perennial grasses. Moderate perennial forbs stature and abundance are being replaced by short-statured annual forbs. Bare ground increases and litter amounts and infiltration rates decline while soil surface temperatures increase. This plant community is resilient, retaining sufficient grazing sensitive native plant species to return to the 1.2 community phases (via Community Phase Pathway 1.3A) by removing prairie dogs and implementing a prescribed grazing system.
Invertebrates: A switch to annual forbs from perennial forbs will not have a significant impact to invertebrates but may reduce season-long nectar producing plants for pollinators. Season-long nectar sources may be found on adjacent plant communities or ecological sites for mobile species. Increased bare ground and prairie dog burrow sites provide increased nesting sites for bumble bees and other ground-nesting insects.
Birds: This very short-statured phase, driven by continued over grazing or prairie dog occupation, is favored by burrowing owls, chestnut-collared longspur, and McCown’s longspur. Prairie dog towns provide abundant prey populations for grassland raptors. The lack of grass and forb stature limits use by many bird species. The invasion of exotic cool-season grasses following Transitional Phase Pathway T1A leads to the 2.0 State and Plant Community Phase 2.3 with no known return pathway due to the presence of exotic cool-season grasses.
Mammals: Suitable food, thermal, shelter, and escape cover (reduction in litter) for most mammals becomes limited. The loss of diversity of grasses and forbs reduces nutrition levels for small and large herbivores, including rodents, white-tailed jackrabbits, and deer. Grazers, such as pronghorn and bison, use prairie dog towns for foraging and loafing.
Amphibians and Reptiles: Prairie dog towns provide habitat for both amphibians and reptiles. Tiger salamanders, prairie rattlesnakes, and other snake species will use the burrow systems of prairie dogs for shelter and denning.
Fish and Mussels: Provides similar life requisites as Community Phase 1.1.
2.0 Native Invaded State
Community Phase 2.1 Western Wheatgrass-Green Needlegrass-Blue Grama: Stabilized:
This plant community develops through Transition pathway T1A, due to changes in management (chronic season-long or heavy late season grazing or complete long-term rest) and the presence of exotic, cool-season grasses. The threshold between states 1.0 and 2.0 is crossed when Kentucky bluegrass, crested wheatgrass, annual bromes, or other exotic species become established. This plant community phase has a very similar appearance and function to the Reference State of Community 1.1, except it has a minor amount of cool-season exotic grasses and forbs. This phase functions at a high level for native wildlife; therefore, managers should consider the 2.0 community phase pathways to avoid transitioning to State 3.0.
Invertebrates: Provides similar life requisites as Community Phase 1.1.
Birds: Provides similar life requisites as Community Phase 1.1.
Mammals: Provides similar life requisites as Community Phase 1.1.
Amphibians and Reptiles: Provides similar life requisites as Community Phase 1.1.
Fish and Mussels: Provides similar life requisites as Community Phase 1.1.
Community Phase 2.2 Blue Grama-Western Wheatgrass/Bare Ground/Exotic Cool-Season Grasses: Active Erosion/Deposition: Blue grama and western wheatgrass will dominate due to naturally occurring active soil erosion/deposition. Continuous grazing or over-utilization will compound the effects of soil erosion. This plant community is 85% grasses and grass-like species, with 10% forbs, and 5% shrubs. The dominant forbs include white sagebrush, pennyroyal, spiny phlox, woolly plantain, scarlet globemallow, and yellow salsify, goldenrod, white heath aster, silverleaf Indian breadroot, and common yarrow. The dominant shrubs are prairie rose, silver sagebrush, western snowberry, and prairie sagewort. However, Wyoming big sage could be present due to the abundance of bare ground and lack of fine fuel to carry a fire. If present, Wyoming big sage will be located along the transition zone between the Badlands landform, not providing sufficient habitat to meet greater sage grouse life requisites. Prescribed grazing with adequate recovery periods (along Community Phase Pathway 2.2A) will help enhance plant vigor and species composition reducing bare ground to regain the cool-season grass and forb diversity components in Community Phase 2.1.
Invertebrates: Provides similar life requisites as Community Phase 1.2.
Birds: Provides similar life requisites as Community Phase 1.2.
Mammals: Provides similar life requisites as Community Phase 1.1.
Amphibians and Reptiles: Provides similar life requisites as Community Phase 1.1.
Fish and Mussels: Provides similar life requisites as Community Phase 1.1.
Community Phase 2.3 Annual Forbs/Bare Ground/Blue Grama/Fetid Marigold (Prairie Dog Town): This plant community is a result of ecological services provided by long-term black-tailed prairie dog occupation coupled with the introduction of exotic cool-season grasses and annual forbs along transitional pathway T1B. Utilizing one or more tools in community phase pathway 2.3A (e.g., removal of black-tailed prairie dogs, control of exotic perennial forbs, implementation of prescribed grazing) can move this community back to phase 2.2; but this may require significant management and economic inputs.
Invertebrates: Provides similar life requisites as Community Phase 1.3.
Birds: Provides similar life requisites as Community Phase 1.3.
Mammals: Provides similar life requisites as Community Phase 1.3.
Amphibians and Reptiles: Provides similar life requisites as Community Phase 1.3.
Fish and Mussels: Provides similar life requisites as Community Phase 1.1.
3.0 Invaded State
Community Phase 3.1 Exotic Cool-Season Grasses/Western Wheatgrass/Forbs: Community phase pathway T2A is characterized by non-use (10 or more years) or low intensity (<20% utilization) grazing and elimination of fire when exotic cool-season grasses are present (as in community phase 2.0). This plant community phase is characterized by a dominance (>30%) to a complete dominance of exotic cool-season grasses (such as Kentucky bluegrass, crested wheatgrass, and/or annual bromes). Restoration pathway R3A requires remnant amounts of native warm-season grasses (i.e., blue grama), cool-season grasses (i.e., needlegrasses, western wheatgrass, prairie Junegrass), and forbs (i.e., silverleaf Indian breadroot and blacksamson echinacea). These remnant populations can only be expressed through frequent prescribed burns and high levels of prescribed grazing management targeting the exotic cool-season grasses. Intensified management along the R3A pathway will have significant short-term negative impacts on wildlife habitat; however, this is necessary to restore long-term native habitat functions.
Invertebrates: Non-use or low intensity (<20% utilization) grazing limits use by beneficial insects provided in States 1.0 and 2.0. Increased litter and lack of grazing leads to limited contact between plant material and mineral soil resulting in a cooler micro-climate, which is unfavorable to most insects. Lack of bare soil limits ground-nesting sites for native bees and other ground- nesting insects. The lack of nectar producing plants limits forage opportunities for bumblebees other pollinating species. This site does not provide life requisites for butterfly species of concern.
Birds: This homogeneous community phase, dominated by exotic plant species, provides limited habitat and life requisites for most obligate grassland-nesting birds. Lack of plant diversity and stature, along with increased litter and the tendency of Kentucky bluegrass and smooth brome to lay down, limits use by many grassland-nesting birds. Litter accumulations reduce use by chestnut-collared and McCown’s longspurs. Woody vegetation invasion is limited to prairie sagewort, not impacting grassland-nesting birds that tend to avoid woody vegetation.
Mammals: Black-tailed prairie dog expansion is possible in this plant community phase. This community phase provides limited foraging habitat for pronghorn and deer. Litter accumulation favors thermal, protective, and escape cover for small rodents. However, reduced availability of native grass seed may reduce food availability for species such as the hispid pocket mouse.
Amphibians and Reptiles: Provides similar life requisites as Community Phase 1.1.
Fish and Mussels: Provides similar life requisites as Community Phase 1.1.
4.0 Invaded Conifer State
Community Phase 4.1 Juniper/Pine:
Resulting from Pathways T2B and T3A, elimination of fire is the major contributor to this community phase crossing the threshold from an herbaceous plant community to a community completely dominated by Rocky Mountain juniper and/or ponderosa pine. Badland Fan sites did not historically support a Rocky Mountain juniper/pine plant community. This phase is dependent upon seed dispersal by birds and mammals from nearby sites. Dense conifers lead to changes in soil chemistry and a change in the associated herbaceous plant community. A detritus layer of juniper needles- leaves, shade, shallow root system, and interception of precipitation with a possible soil chemistry change (decrease in pH) reduces or eliminates an herbaceous or forb understory.
Invertebrates: Junipers are wind-pollinated and thus do not benefit pollinating insects. The loss of a forb component limits insect populations.
Birds: Junipers occur along a continuum. Light infestation may continue to support some grassland and open-area species tolerant of woody invasion (e.g., lark sparrow, vesper sparrow). Bird species intolerant of woody vegetation is eliminated. Species associated with woodlands and woodland edges will increase. The presence of woody plant species may increase predation by mammals and avian predators and brood parasitism by brown- headed cowbirds. Nearly all grassland-nesting bird species are negatively affected by Community Phase 4.1.
Mammals: Nearly all bat species in North Dakota use Community Phase 4.1 for roost sites if mature trees are available. Nearby community phases supporting insects provide foraging opportunities. This phase provides significant thermal, escape and loafing habitat for elk and deer.
Amphibians/Reptiles: Community Phase 4.1 will not support most of the amphibians and reptiles expected to be present in State 1.0, 2.0 and 3.0. Community Phase 4.1 may support some frog species and sagebrush lizards when adjacent to sites supporting sagebrush habitats.
Fish and Mussels: Provides similar life requisites as Community Phase 1.1.
5.0 Go-back State
Community Phase 5.1 Annual/Pioneer-Perennial/Exotic Species: This plant community is the result of severe soil disturbance (such as cropping, recreational activity, or concentrated livestock activity for a prolonged period). Following cessation of disturbances, the resulting plant community is dominated by early pioneer annual and perennial plant species. Plant species composition and production are highly variable. Weedy plants can provide pollinator habitat along with spring and summer cover for many mammals and birds, and their young. Badland Fan ecological sites support short- to mid-statured annual weeds - such as Russian thistle, burning bush (aka kochia), and annual bromes. Russian thistle and burning bush provide seed sources for various grassland-nesting birds and forage for large and small ungulates. They may also provide escape, thermal, and winter cover (depending on location and density).
Successful restoration of native species along transition pathway R5A results in a native grass and forb community in State 2.0. Failed restoration to native species through restoration pathway R5B results in Invaded State 3.0. Wildlife species response will be dependent upon plant community composition, vegetative stature, patch size, and management activities (such as prescribed grazing, burning, inter-seeding, haying, or noxious weed control).
Animal Community -- Grazing Interpretations
This site is well adapted to managed grazing by domestic livestock. The predominance of herbaceous plants across all plant community phases best lends these sites to grazing by cattle; but other domestic grazers with differing diet preferences may also be a consideration depending upon management objectives. Often, the current plant community does not match any particular plant community (as described in the ecological site description). Because of this, a resource inventory is necessary to document plant composition and production. Proper interpretation of this inventory data will permit the establishment of a safe, initial stocking rate for the type and class of animals and level of grazing management. More accurate stocking rate estimates should eventually be calculated using actual stocking rate information and monitoring data.
NRCS defines prescribed grazing as “managing the harvest of vegetation with grazing and/or browsing animals with the intent to achieve specific ecological, economic, and management objectives”. As used in this site description, the term ‘prescribed grazing’ is intended to include multiple grazing management systems (e.g., rotational grazing, twice-over grazing, conservation grazing, targeted grazing, etc.) provided that, whatever management system is implemented, it meets the intent of prescribed grazing definition.
The basic grazing prescription addresses balancing forage demand (quality and quantity) with available forage, varying grazing and deferment periods from year-to-year, matching recovery/deferment periods to growing conditions when pastures are grazed more than once in a growing season, implementation of a contingency (e.g., drought) plan, and a monitoring plan. When the management goal is to facilitate change from one plant community phase or state to another, then the prescription needs to be designed to shift the competitive advantage to favor the native grass and forb species.
Grazing levels are noted within the plant community narratives and pathways in reference to grazing management on adjacent sites. “Degree of utilization” is defined as the proportion of the current years forage production that is consumed and/or destroyed by grazing animals (may refer to a single plant species or a portion or all the vegetation). “Grazing utilization” is classified as slight, moderate, full, close, and severe (see the following table for description of each grazing use category). The following utilization levels are also described in the Ranchers Guide to Grassland Management IV. Utilization levels are determined by using the landscape appearance method as outlined in the Interagency Technical Reference “Utilization Studies and Residual Measurements” 1734-3.
Utilization Level:
Slight (Light) 0-20% Appears practically undisturbed when viewed obliquely. Only choice areas and forage utilized.
Moderate 20-40% Almost all of accessible range shows grazing. Little or no use of poor forage. Little evidence of trailing to grazing.
Full 40-60% All fully accessible areas are grazed. The major sites have key forage species properly utilized (about half taken, half left). Points of concentration with overuse limited to 5 to 10 percent of accessible area.
Close (Heavy) 60-80% All accessible range plainly shows use and major sections closely cropped. Livestock forced to use less desirable forage, considering seasonal preference.
Severe > 80% Key forage species completely used. Low-value forages are dominant.
Hydrological functions
Water is the principal factor limiting herbage production on this site. The site is dominated by soils in hydrologic group B. Infiltration rate is moderate or moderately slow; runoff potential for this site varies from low to very high depending upon surface texture, slope and ground cover. In many cases, areas with greater than 75% ground cover have the greatest potential for high infiltration and lower runoff. An exception would be where short grasses form a dense sod and dominate the site. Areas where ground cover is less than 50% have the greatest potential to have reduced infiltration and higher runoff (refer to Section 4, NRCS National Engineering Handbook for runoff quantities and hydrologic curves).
Recreational uses
The largest acreage of public land available for recreation in the MLRA is owned and managed by the United States Forest Service (USFS) within the Little Missouri National Grasslands in North Dakota (525,211 acres). These areas are available for hunting, fishing, hiking, camping, horse and bike riding, nature viewing, etc. In addition, the Bureau of Land Management (BLM) manages 30,895 acres in North Dakota and Montana with the same recreational opportunities as the USFS lands. North Dakota and Montana Department of Trust Lands (80,220 acres) provide hunting, bird watching, hiking, and other outdoor recreation opportunities. North Dakota Wildlife Management Areas (3,447 acres) of land managed by the states for wildlife habitat in MLRA 58C.
MLRA 58C is home to the North and South Units of Theodore Roosevelt National Park. The Park encompasses approximately 70,000 acres and welcomes approximately 900,000 visitors annually. 29,920 acres of the park is designated Wilderness Area. The south unit of the park has a 48-mile scenic drive while the north unit has a 28-mile scenic drive. The Badland and associated ecological sites provide the main scenery attraction.
Bird watching: Public and private grasslands within MLRA 58C provide essential habitat for prairie-dependent bird species (such as Sprague's pipits, western meadowlark, and Baird's sparrow) along with some of the larger, showy members of the upland prairie including marbled godwits, upland sandpipers, and willets. The abundance of publicly owned lands (such as Theodore Roosevelt National Park, USFS, North Dakota Department of Trust Lands, BLM, etc.) provide excellent birding opportunities. MLRA 58C is in the Central Flyway.
Hunting/Fishing: MLRA 58C is a fall destination for upland game bird hunters, especially sharp-tailed grouse. This MLRA also provides excellent white-tailed deer, mule deer, pronghorn, elk, coyote, and mountain lion hunting opportunities along with the only bighorn sheep hunting units in the North Dakota. The North Dakota Game and Fish Department manages three man-made fishing lakes within the MLRA. Available species include rainbow and brown trout, bluegill, and largemouth and smallmouth bass.
Camping: Many camping opportunities exist in the MLRA. Modern and primitive camping is available at the Theodore Roosevelt National Park, Sully’s Creek State Park, Little Missouri State Park, Buffalo Gap Campground, BLM land, and the Dakota Prairie National Grasslands. The Sully’s Creek and Little Missouri State Parks are designated horse parks.
Hiking/Biking: Over 150 miles of the May-Daah-Hey Trail provide some of the best single-track trails in the world for biking, hiking, or horseback riding. The International Mountain Biking Association (IMBA) has designated the hiking, biking, and horseback riding trail as EPIC - meaning it’s one of the top mountain biking trails in the United States. The trail has nine fenced campgrounds, each accessible by gravel surfaced roads; they include camping spurs, potable water, hitching rails, picnic tables, fire rings, and accessible toilets. They are spaced about every 20 miles along the trail. The North and South Units of the Theodore Roosevelt National Park provide 38.9 and 49.6 miles, respectively, of hiking trails for walkers, bikers, or horseback riders. The Little Missouri State Park has 45 miles of trails that run through the North Dakota Badlands.
Canoeing: Traversing 274 miles through MLRA 58C, the Little Missouri River provides early spring canoeing and kayaking. The Little Missouri River is the only designated State Scenic River in the MLRA. The river passes through Sully Creek State Park, the Little Missouri National Grassland, and Theodore Roosevelt National Park.
Wood products
No appreciable wood products are present on the site.
Other products
Seed harvest of native plant species can provide additional income on this site.
Other information
Site Development and Testing Plan:
Further Investigation is needed on the Sham components included in this ecological site. The central concept of the Badland Fan site is E.C. <8 to a depth of 40 inches. However, many Sham components have been populated E.C. 8-16 within a depth of 16 inches. The soil salinity levels, and associated plant communities needs field documentation. It is currently thought that these components are mostly associated with the Hell Creek formation. Two MLRA map units that have major components of Sham with moderate salinity are:
Patent, gullied, frequently flooded-Sham-Patent complex, 2 to 15 percent slopes (map unit 2q2n9)
Sham loam, 2 to 9 percent slopes (map unit 2qz6q)
This ESD is the best available current knowledge. The site concept and species composition table have been used in the field and tested for more than five years. It is expected that as additional information becomes available revisions may be required.
Supporting information
Inventory data references
Information presented here has been derived from NRCS and other federal/state agency clipping and inventory data. Also, field knowledge of range-trained personnel was used. All descriptions were peer reviewed and/or field-tested by various private, state, and federal agency specialists.
Other references
Abbott, P.L., 2004, Natural disasters, New York, McGraw-Hill Companies, Inc., 460 p.
Bakker, K.K. 2003. The effect of woody vegetation on grassland nesting birds: an annotated bibliography. The Proceedings of the South Dakota Academy of Science 82:119-141.
Barker, W.T. and W. C. Whitman. 1988. Vegetation of the Northern Great Plains. Rangelands 10: 266-272h
Barnhart, Paul. 2017. Documentation of overwintering bat species presence and hibernacula use in the badlands of North Dakota, Northwestern Naturalist 98(1), 48-56, (1 March 2017). https://doi.org/10.1898/NWN15-34.1.
Bjustad, A. J. 1965. Vegetation measurements in relation to range condition classification on the principal range sites of southwestern North Dakota. Ph.D. Thesis. North Dakota State University.
Bluemle. J.P. 2016. North Dakota’s geologic legacy. North Dakota State University Press. 382 pages.
Brand, M. D. and H. Goetz. 1986. Vegetation of exclosures in southwestern North Dakota. Journal of Range Management 39:434-437.
Briske, D.D. (editor). 2017. Rangeland systems – processes, management, and challenges. Springer Series on Environmental Management. 661 pages.
DeKeyser, S., G. Clambey, K. Krabbenhoft and J. Ostendorf. 2009. Are changes in species composition on central North Dakota rangelands due to non-use management? Rangelands 31:16-19.
Dodd, J.L. 1970. Distribution and community site relations of bluebunch wheatgrass in North Dakota. Ph.D. Thesis. North Dakota State University.
Dyke, S. R., S. K. Johnson, and P.T. Isakson. 2015. North Dakota state wildlife action plan – North Dakota Game and Fish Department.
Ehrenfeld, J.G. 2002. Effects of exotic plant invasions on soil nutrient cycling processes. Ecosystems 6:503-523.
Endangered and threatened wildlife and plants; designation of critical habitat for the Dakota skipper and Poweshiek skipperling; Vol. 79 No. Final Rule October 1, 2015, 50 CFR Part 17.
Ereth, C., J. Hendrickson, D. Kirby, E. DeKeyser, K. Sedevic, and M. West. Controlling Kentucky bluegrass with herbicide and burning is influenced by invasion level. Invasive Plant Science and Management 10:80-89.
Flesland, J.R. 1964. Composition and structure of the salt-desert shrub type in the badlands of western North Dakota. M.S. Thesis. North Dakota State University.
Gilgert, W.; and S. Zack. 2010. Integrating multiple ecosystem services introduction ecological site descriptions. Rangelands 32:49-54.
Gillam, Erin. Distribution and habitat use of the bats of North Dakota, Final Report, T2-5-R Bat Survey Final Report 2012_0.pdf (nd.gov).
Grant, T.A. and R.K. Murphy. 2005. Changes on woodland cover on prairie refuges in North Dakota, USA. Natural Areas Journal 25:359-368.
Hanson, H.C and W. Whitman. 1938. Characteristics of major grassland types in western North Dakota. Ecological Monographs 8:57-114.
Heitschmidt, R. K., K. D. Klement, and M. R. Haferkamp. 2005. Interactive effects of drought and grazing on Northern Great Plains rangelands. Rangeland Ecology and Management 58:11-19.
Hendrickson, J.R., P. S. Johnson, M. A. Liebig, K. K. Sedivec, and G. A. Halvorson. 2016. Use of ecological sites in managing wildlife and livestock: an example with prairie dogs. Rangelands 38:23- 28.
Higgins, K.F. 1984. Lightning fires in North Dakota grasslands and in pine-savanna lands of South Dakota and Montana. Journal of Range Management 37:100-103.
Higgins, K.F. 1986. Interpretation and compendium of historical fire accounts in the northern great plains. United States Department of Interior, Fish and Wildlife Service. Resource Publication 161. 39 pages.
Higgins, K. F., A. D. Kruse, and J. L. Piehl. 1987. Effects of fire in the northern Great Plains. South Dakota State University Extension Circular Paper 429.
High Plains Regional Climate Center, University of Nebraska. http://hprcc.unl.edu, Accessed on May 1, 2017.
Hirsch, K.L. 1985. Habitat type classification of grasslands and shrublands of southwestern North Dakota. Ph.D. Thesis. North Dakota State University.
Johnson, S. 2015. Reptiles and amphibians of North Dakota. North Dakota Game and Fish Department. 64 pages.
Jordan, N. R., D.L. Larson, and S.C. Huerd. 2008. Soil modification by invasive plants: effects on native and invasive species of mixed-grass prairies. Biological Invasions 10:177-190.
Mader, E., M. Shepherd, M. Vaughan, and S.H. Black. 2011. Attracting native pollinators: protecting North America's bees and butterflies. Accessed at https://xerces.org, May 1, 2017.
Montana’s State Wildlife Action Plan. 2015. Montana Fish, Wildlife and Parks. Viewed at https://xerces.org/ on May 1, 2017.
North Dakota Division of Tourism, Accessed on February 25, 2019. Available at https://www.ndtourism.com/sports-recreation
North Dakota Parks and Recreation Department, Accessed on February 25, 2019. Available at http://www.parkrec.nd.gov/recreationareas/recreationareas.html
Palit, R., G. and E.S. DeKeyser. 2022. Impacts and drivers of smooth brome (Bromus inermis Leyes.) invasion in native ecosystems. Plants: 10,3390. http://https://www.mdpi.com/2223-7747/11/10/1340
Palit, R., G. Gramig, and E.S. DeKeyser. 2021. Kentucky bluegrass invasion in the Northern Great Plains and prospective management approaches to mitigate its spread. Plants: 10,817. https://doi.org/10.3390/plants10040817
Printz, J.L. and J.R. Hendrickson. 2015. Impacts of Kentucky bluegrass invasion (Poa pratensis) on ecological processes in the Northern Great Plains. Rangelands 37:226-232.
Redmann, R.E. 1975. Production ecology of grassland plant communities in western North Dakota. Ecological Monographs 45:83-106.
Reeves, J.L., J.D. Derner, M.A. Sanderson, J.R. Hendrickson, S.L. Kronberg, M.K. Petersen, and L.T. Vermeire. 2014. Seasonal weather influences on yearling beef steer production in C3-dominated Northern Great Plains rangeland. Agriculture, Ecosystems and Environment 183:110-117.
Robinson, A.C. 2014. Management plan and conservation strategies for greater sage grouse in North Dakota. North Dakota Game and Fish Department.
Royer, R. A., 2003. Butterflies of North Dakota: an atlas and guide. Minot State University.
Sanford, R.C. 1970. Skunk bush in the North Dakota badlands: ecology, phytosociology, browse production, and utilization. Ph.D. Thesis. North Dakota State University.
Seabloom, R. 2020. Mammals of North Dakota. North Dakota State University Press.
Sedivec, K.K., and J.L. Printz. 2014. Ranchers guide to grassland management IV. North Dakota State University Extension Service publication R1707.
South Dakota Dept. of Game, Fish and Parks. 2014. South Dakota wildlife action plan. Wildlife Division Report 2014-03.
Spaeth, K.E., Hayek, M.A., Toledo, D., and Hendrickson, J. 2019. Cool season grass impacts on native mixed grass prairie species in the Northern Great Plains. America’s Grassland Conference: Working Across Boundaries. The Fifth Biennial Conference on the Conservation of America’s Grasslands. Bismarck, ND. 20-22 August.
Steffens, Tim, G. Grisson, M. Barnes, F. Provenza, and R. Roath. Adaptive grazing management for recovery. Know why you’re moving from paddock to paddock. Rangelands 35(5):28–34
Tidwell, D., D.T. Fogarty, and J.R. Weir. 2021. Woody encroachment in grasslands, a guide for understanding risk and vulnerability. Oklahoma State University, Oklahoma Cooperative Extension Service Publication E-1054. 32 pages.
Toledo, D., M. Sanderson, K. Spaeth, J. Hendrickson, and J. Printz. 2014. Extent of Kentucky bluegrass and its effect on native plant species diversity and ecosystem services in the Northern Great Plains of the United State. Invasive Plant Science and Management 7:543-552.
United States Department of Agriculture, Natural Resources Conservation Service. 2006. Land Resource Regions and Major Land Resource Areas of the United States, the Caribbean, and the Pacific Basin. U.S. Department of Agriculture Handbook 296.
USDA, NRCS. National Water and Climate Center. (http://www.wcc.nrcs.usda.gov) USDA, NRCS. National range and pasture handbook, September 1997.
USDA, NRCS. National Soil Information System, Information Technology Center. Accessed on May 1, 2017 at http://nasis.nrcs.usda.gov.
USDA, NRCS. 2001. The PLANTS Database, Version 3.1. http://plants.usda.gov. Accessed May 2, 2017.
USDA, NRCS. 2021. Species diversification of crested wheatgrass dominated grasslands: a review of methods. Plant Materials Technical Note No. MT-126 November 2021.
USDA, NRCS, Various published soil surveys.
USDI BLM. Utilization studies and residual measurements. Interagency Technical Reference 1734-3. 1999.
Vinton, M.A. and E.M. Goergen. 2006. Plant-soil feedbacks contribute to the persistence of Bromus inermis in tallgrass prairie. Ecosystems 9:967-976.
Whitman, W., H. Hanson, and R. Peterson. 1943. Relation of drought and grazing to North Dakota rangelands. North Dakota Agricultural Experimentation Bulletin 320.
Zaczkowski, N. K. 1972. Vascular flora of Billings, Bowman, Golden Valley, and Slope counties, North Dakota. Ph.D. Thesis, North Dakota State University.
Zimmerman, G. M. 1981. Effects of fire upon selected plant communities in the little Missouri badlands. M.S. Thesis, North Dakota State University.
Contributors
ND NRCS: David Dewald, Alan Gulsvig, Mark Hayek, Jeanne Heilig, John Kempenich, Chuck Lura, Jeff Printz, and Steve Sieler.
Approval
Suzanne Mayne-Kinney, 4/21/2025
Acknowledgments
NRCS would like to acknowledge the United State Forest Service (USFS) and National Park Service (NPS) for access to USFS and NPS properties and technical assistance in ESD development. USFS: Jack Dahl, Nickole Dahl, Chad Prosser, Jack Butler; NPS: Chad Sexton.
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Rangeland health reference sheet
Interpreting Indicators of Rangeland Health is a qualitative assessment protocol used to determine ecosystem condition based on benchmark characteristics described in the Reference Sheet. A suite of 17 (or more) indicators are typically considered in an assessment. The ecological site(s) representative of an assessment location must be known prior to applying the protocol and must be verified based on soils and climate. Current plant community cannot be used to identify the ecological site.
Author(s)/participant(s) | USDA NRCS North Dakota |
---|---|
Contact for lead author | NRCS State Rangeland Management Specialist |
Date | 04/21/2025 |
Approved by | Suzanne Mayne-Kinney |
Approval date | |
Composition (Indicators 10 and 12) based on | Annual Production |
Indicators
-
Number and extent of rills:
Rills are not expected on this site in PCP (Plant Community Phase) 1.1 except after multi-year drought. In PCP 1.2, rills will be short (less than 16 inches) and disconnected. -
Presence of water flow patterns:
In PCP 1.1, water flow patterns are not apparent. In PCP 1.2, water flow patterns are visible, long (greater than 10 feet) and disconnected. -
Number and height of erosional pedestals or terracettes:
Neither pedestals nor terracettes are expected in PCP 1.1. In PCP 1.2, pedestaled plants are common with terracettes also visible. -
Bare ground from Ecological Site Description or other studies (rock, litter, lichen, moss, plant canopy are not bare ground):
In PCP 1.1, bare ground ranges from 5 to 15%; patch size is 2 inches or less in diameter and disconnected. Animal activity (burrows and ant mounds) may result in infrequent bare ground patches of up to 24 inches in diameter. In PCP 1.2, bare ground ranges from 50 to 65%; patch size ranges from 12 to 24 inches with many of the bare ground patches connected. -
Number of gullies and erosion associated with gullies:
Active gullies are not expected in PCP 1.1. If present, gully channel(s) are fully vegetated with no active erosion visible. Active gullies are expected in PCP 1.2. Gullies in PCP 1.2 will be greater than 10 feet in length and have signs of active erosion. -
Extent of wind scoured, blowouts and/or depositional areas:
Wind-scoured areas or depositional areas do not occur on this site. -
Amount of litter movement (describe size and distance expected to travel):
Plant litter movement is not expected in PCP 1.1. Plant litter movement is easily visible in PCP 1.2 with small/fine class of plant litter moving up to 3 feet. -
Soil surface (top few mm) resistance to erosion (stability values are averages - most sites will show a range of values):
Stability class averages 4 or greater in PCP 1.1. Stability class averages 2 or greater in PCP 1.2. -
Soil surface structure and SOM content (include type of structure and A-horizon color and thickness):
PCP 1.1: Primary surface structure is fine granular. A-horizon is 1 to 4 inches thick with very dark grayish brown (2.5Y 3/2 moist) or grayish brown (2.5Y 5/2 dry) colors. PCP 1.2: Primary surface structure is thin platy. A-horizon is less than 1 inch thick with dark grayish brown (2.5Y 4/2 moist) or light brownish gray (2.5Y 6/2 dry) colors. -
Effect of community phase composition (relative proportion of different functional groups) and spatial distribution on infiltration and runoff:
In PCP 1.1, mid- and short statured bunchgrasses are dominant and well distributed across the site. Mid- and short-statured rhizomatous grasses and a diverse forbs component are subdominant. In PCP 1.2, mid- and short-statured warm-season bunchgrasses and grass-likes are dominant with mid- and short-statured rhizomatous grasses, forbs, and shrubs are subdominant. -
Presence and thickness of compaction layer (usually none; describe soil profile features which may be mistaken for compaction on this site):
No compaction layers occur naturally on this site. In PCP 1.2, some platy soil structure may be noted but this is due to soil deposition, not compaction. -
Functional/Structural Groups (list in order of descending dominance by above-ground annual-production or live foliar cover using symbols: >>, >, = to indicate much greater than, greater than, and equal to):
Dominant:
Phase 1.1:
Mid & short C4 bunch grasses (4); Mid & short C3 bunch grasses (4
Phase 1.2:
Mid & short C4 bunch grasses (4); Grass-likes (2)Sub-dominant:
Phase 1.1:
Mid & short C3 rhizomatous grasses (1); Forbs (6)
Phase 1.2:
Mid & short C3 rhizomatous grasses (1); Forbs (6); Mid & short C4 rhizomatous grasses (1); Shrub (4)Other:
Minor
Phase 1.1:
Mid & short C4 rhizomatous grasses; Grass-likes; Shrub
Minor - Phase 1.2:
Mid & short C3 bunchgrassesAdditional:
Due to differences in phenology, root morphology, soil biology relationships, and nutrient cycling Kentucky bluegrass, smooth brome, and crested wheatgrass are included in a new Functional/structural group, mid- and short-statured early cool-season grasses (MSeC3), not expected for this site.
To see a full version 5 rangeland health worksheet with functional/structural group tables, please use the following hyperlink:
https://efotg.sc.egov.usda.gov/references/public/ND/58C_Badland_Fan_Narrative_Final_Ref_FSG.pdf -
Amount of plant mortality and decadence (include which functional groups are expected to show mortality or decadence):
Not expected in PCP 1.1. Some dead or dying plants/plant parts would be present in PCP 1.2, generally associated with pedestalled plants and/or gullies. -
Average percent litter cover (%) and depth ( in):
Plant litter cover in PCP 1.1 is 50 to 75%, 0.25 5o 0.5 inches thick, and in contact with soil surface. In PCP 1.2, plant litter cover ranges from 1 to 30%, less than 0.25 inches thick and in contact with soil surface. -
Expected annual annual-production (this is TOTAL above-ground annual-production, not just forage annual-production):
Annual air-dry production for PCP 1.1 is 1200 lbs./ac (reference value) with normal precipitation and temperatures. Low and high production years should yield 800 lbs./ac to 1600 lbs./ac, respectively. Annual air-dry production for PCP 1.2 is 600 lbs./ac (reference value) with normal precipitation and temperatures. Low and high production years should yield 400 lbs./ac to 800 lbs./ac, respectively. -
Potential invasive (including noxious) species (native and non-native). List species which BOTH characterize degraded states and have the potential to become a dominant or co-dominant species on the ecological site if their future establishment and growth is not actively controlled by management interventions. Species that become dominant for only one to several years (e.g., short-term response to drought or wildfire) are not invasive plants. Note that unlike other indicators, we are describing what is NOT expected in the reference state for the ecological site:
State and local noxious species, Kentucky bluegrass, smooth bromegrass, crested wheatgrass, and Rocky Mountain and prostrate juniper. -
Perennial plant reproductive capability:
In PCP 1.1, noninvasive species in all functional/structural groups are vigorous and capable of reproducing annually under normal weather conditions. In PCP 1.2, some reduced vigor and reproductive capability would be expected.
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The Ecosystem Dynamics Interpretive Tool is an information system framework developed by the USDA-ARS Jornada Experimental Range, USDA Natural Resources Conservation Service, and New Mexico State University.
Click on box and path labels to scroll to the respective text.
Ecosystem states
States 2 and 5 (additional transitions)
T1A | - | Introduction of exotic cool-season grasses |
---|---|---|
T2A | - | Extended periods of non-use or very light grazing, no fire |
T2B | - | Extended periods of non-use or very light grazing, no fire |
R3A | - | Long-term prescribed grazing and prescribed burning with possible range planting |
T3A | - | Extended periods of non-use or very light grazing, no fire |
R4A | - | Prescribed burning and/or chemical/mechanical brush management |
R4B | - | Prescribed burning and/or chemical/mechanical brush management |
R5A | - | Successful range planting with prescribed grazing and prescribed burning |
R5B | - | Failed range planting and/or secondary succession |
T6A | - | Elimination of annual cropping |
State 1 submodel, plant communities
1.1A | - | Soil erosion and deposition |
---|---|---|
1.1B | - | Long-term occupation by prairie dogs |
1.2A | - | Soil stabilization |
1.2B | - | Long-term occupation by prairie dogs |
1.3A | - | Removal of prairie dogs |
State 2 submodel, plant communities
2.1A | - | Soil erosion and deposition |
---|---|---|
2.1B | - | Long-term prairie dog occupation |
2.2A | - | Soil stabilization |
2.3A | - | Prairie dog abandonment/removal |