Physiographic features

This site typically occurs on sandy uplands – sand plains, delta plains, and lake plains. It occurs on flats and on foot slopes of rises. A few areas are on terraces. The parent materials are glaciofluvial or sandy glaciolacustrine deposits. Slopes range from 0 to 6 percent.

Landform: sand plain, delta plain, lake plain, terrace

Table 2. Representative physiographic features

Climatic features

MLRA 55B is considered to have a continental climate – cold winters and hot summers, low humidity, light rainfall, and much sunshine. Extremes in temperature are characteristic of the MLRA. The climate is the result of this MLRA’s location in the geographic center of North America. There are few natural barriers on the northern Great Plains. The air masses move unobstructed across the plains and account for rapid changes in temperature.

Annual precipitation ranges from 18 to 23 inches per year. The normal average annual temperature is about 41.5° F. January is the coldest month with average low temperature ranging from about -4.3° F (Petersburg, ND) to about 2.5° F (Mellette, SD). July is the warmest month with temperatures averaging from about 79° F (Petersburg, ND) to about 84° F (Mellette, SD). The range of normal average monthly temperatures between the coldest and warmest months is about 64° F. This large annual range attests to the continental nature of this MLRA's climate. Winds average about 11 miles per hour annually, ranging from about 13 miles per hour during the spring to about 10 miles per hour during the summer. Daytime winds are generally stronger than nighttime and strong storms may bring brief periods of high winds with gusts to more than 50 miles per hour.

Growth of native cool-season plants begins in late March and continues to early to mid-July. Native warm- season plants begin growth in mid-May and continue to the end of August. Greening up of cool-season plants can occur in September and October when adequate soil moisture is present.

Table 3. Representative climatic features

Climate stations used

• (1) BUTTE 5SE [USC00321225], Butte, ND

• (2) CARRINGTON [USC00321360], Carrington, ND

• (3) FORMAN 5 SSE [USC00323117], Forman, ND

• (4) HARVEY 4NE [USC00324013], Harvey, ND

• (5) LA MOURE [USC00324937], Lamoure, ND

• (6) MELLETTE 4 W [USC00395456], Northville, SD

• (7) PETERSBURG 2 N [USC00327027], Petersburg, ND

• (8) COLUMBIA 8 N [USC00391873], Columbia, SD

Influencing water features

This site does not receive significant additional water, either as runoff from adjacent slopes or from a seasonal high-water table. Although the seasonal water table can be as shallow as 1.5 feet early in the growing season on low-relief areas, the claypan layer, which affects root growth, prohibits the plants from benefiting significantly from subirrigation. Depth to the water table typically is greater than 4 feet through most of the growing season. Surface infiltration is moderately rapid or moderate. Permeability is moderately rapid or rapid in the surface soil, but slow to moderate in the claypan subsoil. Water loss is through percolation below the root zone and through evapotranspiration.

Soil features

Soils associated with Sandy Claypan ecological site are in the Mollisol order, which are classified further as Calcic Natrudolls. These soils were developed under prairie vegetation. Salt accumulations, where present, are below a depth of 16 inches. They formed in glaciofluvial or glaciolacustrine deposits; in some soils, glacial till occurs below a depth of 4 feet.

The common feature of soils in this site is a sodic, claypan subsoil layer that is fine sandy loam, sandy loam, or loam (with <18% clay); it forms a ribbon <1 inch long. Although the soil parent materials are very deep, the claypan layer, which affects root growth, occurs in the upper part of the subsoil (at a depth of 6 to 20 inches). The texture of the surface layer is typically fine sandy loam or sandy loam, but loam and loamy fine sand are included. These soils commonly have a gray subsurface layer of loamy find sand just above the claypan layer. Salt accumulations, where present, are below a depth of 16 inches. The soils in this site typically are moderately well drained – redoximorphic features, where present, are deeper than 1.5 feet.

Soil salinity is typically none to very slight above the claypan layer (E.C. <4 dS/m); however, slight salinity (E.C. 4-8 dS/m) is allowable within a depth of 16 inches. Below 16 inches, it may increase to moderate (E.C. 8 - <16 dS/m) in some soils. Sodicity is low (SAR <5) above the claypan layer; but increases to moderate or moderately high (SAR 5-15) in the subsoil and substratum. Soil reaction is strongly acid to slightly alkaline (pH 5.1 to 7.8) above the claypan and slightly alkaline to strongly alkaline (pH 7.4 to 9.0) in the subsoil and substratum. Calcium carbonate content is typically none in the surface soil and upper few inches of the claypan layer; below this, it may increase to moderate (5-15% CaCO3).

These soils are mainly susceptible to wind erosion. Loss of the soil surface layer can result in a shift in species composition and/or production.

The major soil series correlated to Sandy Claypan site is Letcher.

Access Web Soil Survey ( https://websoilsurvey.sc.egov.usda.gov/App/WebSoilSurvey.aspx ) for specific local soils information.

Table 4. Representative soil features

Animal community

Animal Community – Wildlife Interpretations

Landscape

The MLRA 55B landscape is characterized by mostly nearly level to gently rolling till plains with some steep slopes adjacent to streams and many poorly defined drainage channels. The continental drainage divide occurs in the east central part of the MLRA. The MLRA is located within the Prairie Pothole Region with temporary, seasonal, and semi-permanent wetlands throughout the MLRA. The MLRA includes areas of eskers, kames, and ground moraines. MLRA 55B is considered to have a continental climate with cold winters and hot summers, low humidity, light rainfall, and much sunshine. Extremes in temperature are common and characteristic of the MLRA. This area supports mid- to tall-grass prairie vegetation with American elm, bur oak, green ash, and willow species growing along the riparian zones of river systems found throughout the MLRA. Complex intermingled ecological sites create diverse grass/shrub land habitats interspersed with varying densities of linear, slope, depressional, and in-stream wetlands associated with headwater streams and tributaries of the James, Pipestem, Maple, Goose, Sheyenne, Wild Rice, and Elm Rivers. MLRA 55B is located within North and South Dakota and within the boundaries of the Prairie Pothole Region.

Three Hydrologic Unit Areas make up this MLRA. Approximately 6% drains into the Mouse River into MLRA 55A, with the balance split between the James and Sheyenne Rivers.

By the mid-19th century, over 76% of the MLRA had been converted from mid- to tall-grass prairie to annual crop production. To alleviate crop production loss from wetlands and overland flow, a system of shallow surface ditches, judicial ditches, and road ditches removes surface water in spring and during high rainfall events. Tile drainage systems have been or are being installed extensively throughout MLRA 55B for sub-surface field drainage to enhance annual crop production.

Historic Communities/Conditions within MLRA 55B:

The northern tall- and mixed-grass prairie were disturbance-driven ecosystems with fire, herbivory, and climate functions as the primary ecological drivers (either singly or often in combination). American bison roamed MLRA 55B, wintering along the Mouse River in MLRA 55A and migrating through MLRA 55B and into MLRA 56A. Many species of grassland birds, small mammals, insects, reptiles, amphibians, elk, moose, pronghorn, white-tailed deer, and large herds of American bison were historically among the inhabitants adapted to this region. Roaming herbivores, as well as several small mammals and insect species, were the primary consumers linking the grassland resources to large predators (such as the wolf, American black bear, grizzly bear) and smaller carnivores (such as the coyote, bobcat, red fox, and raptors). Extirpated species include free- ranging American bison and gray wolf (breeding). Extinct is the Rocky Mountain locust.

Present Communities/Conditions within MLRA 55B:

This area supports natural prairie vegetation characterized by western wheatgrass, green needlegrass, needle and thread, and blue grama. Little bluestem is an important species on the more sloping and shallower soils. Prairie cordgrass, northern reedgrass, big bluestem, and wheat sedge (aka slough sedge) are important species on wet soils. Western snowberry, leadplant, and prairie rose are commonly interspersed throughout the area.

Over 80% of MLRA 55B has been converted to annual crop production. These influences fragmented the landscape, reduced or eliminated ecological drivers (fire), and introduced exotic plant species including smooth brome, crested wheatgrass, Kentucky bluegrass, and leafy spurge; this further impacted plant and animal communities. The loss of the bison and fire as primary ecological drivers greatly influenced the character of the remaining native plant communities and the associated wildlife, moving towards a less diverse and more homogeneous landscape. Annual cropping is the main factor contributing to habitat fragmentation, reducing habitat quality for area- sensitive species.

Hydrological manipulation is extensive throughout the MLRA. Extensive wetland and subsurface tile drainage have taken place. Straightened segments of ephemeral and intermittent tributary streams of the James, Wild Rice, and Sheyenne River have reduced sinuosity, created oxbows, and enabled the conversion of riparian ecological sites to annual crop production. These anthropogenic impacts have reduced flood water detention and retention on the landscape. The results have been increasing storm water runoff sediment and nutrient loading to the James and Sheyenne Rivers and their tributaries (along with lakes and reservoirs within the MLRA). Large dams on the James, Pipestem and Sheyenne rivers, along with installation of instream structures have reduced aquatic species movement within the MLRA.

National wildlife refuges, waterfowl production areas, state wildlife management areas, and North and South Dakota Department of Trust Lands provide herbaceous and woody cover for wildlife. In addition, the United States Army Corps of Engineers and the United States Bureau of Reclamation manage three man-made reservoirs - Jamestown Reservoir, Pipestem Reservoir, and Lake Ashtabula for flood control, also providing fish habitat and adjacent uplands for wildlife cover. Lonetree Wildlife Management Area (WMA) is the largest state managed wildlife area covering 32,800 acres. Arrowwood National Wildlife Refuge is the largest refuge consisting of 16,000 acres.

Characteristic wildlife species in this area are:

Birds: Common goldeye, bufflehead, broad-winged hawk, alder flycatcher, mourning warbler, mallard, blue-winged teal, red-tailed hawk, American kestrel, killdeer, eastern and western kingbird, western meadowlark, American crow, common yellowthroat, clay-colored sparrow, vesper sparrow, red-necked grebe, Savannah sparrow, downy and hairy woodpeckers, black-capped chickadee, white-breasted nuthatch, and brown-headed cowbird.

Mammals: Northern short-tailed shrew, white-tailed jackrabbit, Franklin’s ground squirrel, thirteen- lined ground squirrel, northern pocket gopher, plains pocket gopher, western harvest mouse, deer mouse, meadow vole, meadow jumping mouse, western jumping mouse, coyote, red fox, racoon, American badger, striped skunk, white-tailed deer, elk, moose, beaver, muskrat, mink, weasel, woodchuck, and red, eastern gray and fox squirrels.

Reptiles/Amphibians: American toad, Great Plains toad, northern leopard frog, chorus frog, tiger salamander, plains garter snake, smooth green snake, wood frog, and common garter snake.

Presence of wildlife species is often determined by ecological site characteristics including grass and forb species, tree and shrub species, hydrology, aspect, and other associated ecological sites. The home ranges of a majority of species are usually larger than one ecological site or are dependent upon more than one ecological site for annual life requisites. Ecological sites offer different habitat elements as the annual life requisites change. Habitat improvement and creation must be conducted within the mobility limits of a known population for the species.

Insects play an important role in providing ecological services for plant community development. Insects that are scavengers or aid in decomposition provide the food chain baseline sustaining the carnivorous insects feeding upon them. Many insects provide the ecological services necessary for pollination, keeping plant communities healthy and productive. Insects provide a protein food source for numerous species including grassland-nesting birds, woodpeckers, woodland edge and interior species, and their young. Extensive use of insecticides for specialty crops such as soybeans, corn, and other crops has greatly reduced insects within this MLRA.

Species of Concern within MLRA 55B:

The following is a list of species considered “species of conservation priority” in the North Dakota State Wildlife Action Plan (2015) and South Dakota State Wildlife Action Plan (2014); and species listed as “threatened, endangered, or petitioned” under the Endangered Species Act within MLRA 55B at the time this section was developed:

Invertebrates: Dakota skipper, Iowa skipper, monarch butterfly, northern sandy tiger beetle, Ottoe skipper, Poweshiek skipperling, regal fritillary, yellow-banded bumble bee, and western bumble bee.

Birds: America avocet, American bittern, American kestrel, American white pelican, Baird’s sparrow, bald eagle, black-billed cuckoo, black tern, bobolink, burrowing owl, canvasback, chestnut-collared longspur, Dickcissel, ferruginous hawk, Franklin’s gull, grasshopper sparrow, horned grebe, lark bunting, LeConte’s sparrow, lesser scaup, marbled godwit, Nelson’s sparrow, northern goshawk, northern harrier, northern pintail, osprey (migration), peregrine falcon (migration), piping plover (migration), red knot (migration), sharp-tailed grouse, short-eared owl, Swainson’s hawk, upland sandpiper, western meadowlark, willet, Wilson’s phalarope, whooping crane (migration), and yellow rail.

Mammals: Arctic shrew, big and little brown bats, Franklin’s ground squirrel, northern river otter, plains pocket mouse, pygmy shrew, Richardson’s ground squirrel, and silver-haired bat.

Amphibians and Reptiles: Canadian toad, plains hognose snake, smooth green snake, and snapping turtle.

Fish and Mussels: Black sandshell, blacknose shiner, Carmine shiner, creek heelsplitter, creeper, deertoe, fragile papershell, mapleleaf, northern pearl dace, northern redbelly dace, pink heelsplitter, threeridge, trout-perch, yellow sandshell, and Wabash pigtoe.

Grassland Management for Wildlife in MLRA 55B

Management activities within the community phase pathways impact wildlife but are essential for maintenance of healthy grassland ecosystems. Community phase, transitional, and restoration pathways are keys to long-term management within each State and between States. Timing, intensity, and frequency of these inputs can have dramatic positive or negative effects on local wildlife species. Ranchers and other land managers must always consider the long-term beneficial management effects of grassland and woodland resources in comparison to typically short-term negative effects to the habitats of individual species.

Ecological sites occur as intermingled complexes on the landscape with gradual or sometimes abrupt transitions. Rarely do ecological sites exist in large enough acreage to manage independently. Ecological sites supporting a dominance of herbaceous vegetation (Loamy/Sandy) can be located adjacent to ecological sites that support medium to tall shrubs (Loamy Overflow or Loamy-Wooded State). Conversely, ecological sites that are dominated by short- to mid-statured grasses (Claypan) can be adjacent to sites with bare soil only supporting minor amounts of short grasses and forbs (Thin Claypan).

Management of these complex ecological sites can provide a heterogeneous or a homogenous landscape. Grassland bird use reduces as the plant community transitions to a homogenous state. Managers must recognize ecological sites and the complexes in which they occur to properly manage the landscape. A management regime for one ecological site may negatively impact an adjacent site, e.g., alteration of a grazing regime within the Invaded Wooded State of Loamy ecological site to encourage understory growth may encourage exotic cool-season grasses to increase or dominate an adjacent ecological site.

Life requisites and habitat deficiencies are determined for targeted species. Deficiencies need to be addressed along community phase, transitional, and restoration pathways as presented in specific state-and-transition models. Ecological sites should be managed and restored within the site’s capabilities to provide sustainable habitat for targeted species or species guilds. Managers also need to consider vegetative associations provided by adjacent/intermingled ecological sites for species with home ranges or life requisites that may not be provided by one ecological site.

Grassland-nesting birds use various grass heights for breeding, nesting, foraging, or winter habitat. While most species use varying heights, many have a preferred vegetative stature height or sensitivity to woody vegetation. Understanding the sensitivity of grassland species to woody vegetation and preferred vegetative structure enables managers to determine which grassland-nesting bird species avoid grassland habitats adjacent to woody dominated plant community. The following chart provides sensitivity to woody vegetation and preferred vegetative stature heights.

To see the chart please follow the hyperlink:
https://efotg.sc.egov.usda.gov/references/public/ND/55B_Sandy_Claypan_Narrative_FINAL_Ref_FSG.pdf

Sandy Wildlife Habitat Interpretation

Sandy Claypan ecological sites have a sodic, claypan subsoil layer which occurs at a depth of 6 to 20 inches. Sandy Claypan sites support tall-warm season species but become dominated by short- warm season grasses caused by drought or heavy grazing pressure. Sandy Claypan sites support diverse stands of tall and short warm-season grasses, along with a diverse stand of cool-season grass and numerous forb species. Associated ecological sites include Limy Subirrigated, Saline Lowland, Sands, Sandy, Subirrigated Sands, Sodic Subirrigated, and Thin Claypan. This complex of ecological sites provides habitat for many edge-sensitive grassland bird species.

Sandy Claypan habitat features and components commonly support grassland-nesting birds, notably nesting and brood cover and lekking sites for sharp-tailed grouse, dependent upon its state. Insects rely on associated forbs and grasses for survival; the insects serve as food sources for birds and their young.

Sandy Claypan ecological sites may be found in four plant community states (1.0 Reference State, 2.0 Native/Invaded State, 3.0 Invaded State, and 4.0 Go-back State) within a local landscape. Multiple plant community phases exist within each state. Today, these states occur primarily in response to grazing and drought. Secondary influences include anthropogenic disturbances, black- tailed prairie dogs, and fire.

Because there is no known restoration pathway from State 2.0 to State 1.0, it is important to intensively manage using tools in State 1.0 and State 2.0 community phase pathways to prevent further plant community degradation along the T1A transitional pathway to Native/Invaded State 2.0 to 3.0 Native Invaded State thresholds. Native wildlife generally benefits from a heterogeneous grassland found in States 1.0 and 2.0 that include diverse grass and forb species with varying structure and density. As plant communities degrade within State 2.0, short warm-season grasses increase while native forbs are reduced. This transition results in reduced structure and increased plant community homogeneity. When adjacent/intermingled ecological sites undergo the same transition, the result can be an expansive, homogenous landscape.

State 3.0 occurs when cool-season exotic grasses, such as crested wheatgrass, begin to dominate the site. Reduced forb diversity limits insect populations, negatively affecting grassland nesting bird foraging opportunities. Increased lesser spikemoss can limit access to bare ground by nesting. A homogenous grassland landscape does not provide quality escape or winter cover. As a result, many species are not able to meet life requisites. State 3.0 has dramatic increases of short, warm- season grasses and lesser spikemoss with a further reduction in native forbs. Successful restoration along the R3A pathway with native vegetation seeding and mechanical treatment will be expensive and dependent upon climatic and seedbed conditions.

Likewise, success along restoration pathways R3A and R4A from State 3.0 and 4.0 to State 2.0 is very difficult and is dependent upon presence of a remnant native grass population. This concept also applies to wildlife as the target species must either be present on adjacent State 1.0 or State 2.0 plant communities or on ecological sites within the species’ mobility limits. Species with limited mobility, such as Dakota skippers, must exist near the plant community to utilize restored sites.
Mobile species (such as grassland nesting birds) can easily locate isolated, restored plant communities.

Management along community phase, transition or restoration pathways should focus upon attainable changes. Short- and long-term monetary costs must be evaluated against short- and long- term ecological services in creating and maintaining habitat of sufficient quality to support a sustainable population density.

1.0 Reference State

Community Phase 1.1 Prairie Sandreed-Bluestems-Needlegrasses-Wheatgrasses: This plant community offers quality wildlife habitat; every effort should be made to maintain this ecological site within this community phase. This phase retains high functionality through continued maintenance, including prescribed grazing with adequate recovery period as well as prescribed fire. Predominance of grass species in this community favors grazers and mixed-feeders (animals selecting grasses as well as forbs and shrubs). The structural diversity provides habitat for a wide array of migratory and resident birds.

Invertebrates: Insects play a role in maintaining the forb community and provide a forage base for grassland birds, reptiles and rodents. Ecological services, historically provided by bison are mirrored by domestic livestock. These services include putting plant material and dung in contact with mineral soil to be used by low trophic level consumers (such as invertebrate shredders, predators, herbivores, dung beetles, and fungal-feeders).

This ecological site and plant community may be used by Dakota skippers since host plants, such as little bluestem and sand dropseed, can be present. Regal fritillary habitat is limited due to the rarity of Nuttall's violet and prairie violets. Monarch butterfly may use flowering forbs on this site; however, few milkweed species are found on this site to support breeding. Bumblebees and other native bees utilize forbs as a nectar source and bare ground for nesting sites in bunchgrasses. Prescribed grazing with adequate recovery periods, as well as prescribed fire, to maintain the 1.1 phase has little effect on nests of ground- dwelling insects.

Birds: This plant community provides quality nesting, foraging, and escape habitats favored by mid- to tallgrass-nesting birds. Several species of grassland birds that prefer mid- to tallgrass stature will use this site. In years with reduced precipitation or drought, nesting recruitment may be compromised. This plant community provides suitable areas for sharp-tailed grouse nesting and brood-rearing habitat. Diverse prey populations provide good hunting opportunities for grassland raptors.

Mammals: The diversity of grasses and forbs provide high nutrition levels for small and large herbivores including voles, mice, rodents, jackrabbits, pronghorn, and deer. Short to moderate stature provides suitable food, thermal, protective, and escape cover for small herbivores.

Amphibians/Reptiles: The Sandy Claypan ecological site and associated plant communities provide habitat for smooth green snakes, plains hog-nosed snake (prefer sandy soils), and northern prairie skink (secondary range in MLRA 55B). Habitat for the northern leopard frog and Canadian toad is dependent upon distance to water features (such as wetlands, lakes, or streams).

Fish and Mussels: This ecological site is not directly associated with streams, rivers, or water bodies. Associated ecological sites, do not provide significant additional water to this site. Management on these interconnected sites will have limited, secondary effects on aquatic species.

Community Phase 1.2 Blue Grama/Sedges/Western Wheatgrass: Multi-year drought, with or without heavy grazing pressure, increases the percentage of blue grama and sedges in this plant community. This plant community becomes dominated by short, warm-season grasses, changing the stature of plant community from mid- to tall-grasses to mid- to short-grass species.

Invertebrates: Multi-year drought may negatively impact ground-nesting sites for bumblebees, other native bees, and other ground-nesting insects due to reduction of forbs, timing or lack of forb flowering, or increased soil compaction during wet conditions.

Birds: This plant community provides quality nesting, foraging, and escape habitats favored by shortgrass- nesting birds. A shift to a shorter plant structure (along Community Phase Pathway 1.1A) benefits McCown’s longspur, chestnut collared longspur, horned lark, and burrowing owl. Species that prefer a midgrass stature may be successful with normal to above-normal precipitation and a change in management along the 1.2A Community Phase Pathway. Sharp-tailed grouse may still use this plant community for leks and brood rearing; however, winter cover must be provided by adjacent ecological sites or plant communities. Limited cover and diverse prey populations provide good hunting opportunities for grassland raptors.

Mammals: Shorter statured grasses reduce thermal, escape, and protective cover for mammals.

Amphibians/Reptiles: Provides similar life requisites as Community Phase 1.1.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

2.0 Native/Invaded State

Community Phase 2.1 Prairie Sandreed-Bluestems-Needlegrasses-Wheatgrasses: This plant community develops through Transition Pathway T1A, due to changes in management (heavy season-long grazing or complete rest) and the presence of exotic, cool-season grasses. The threshold between States 1.0 and 2.0 is crossed when Kentucky bluegrass, crested wheatgrass, smooth brome, or other exotic species become established. This plant community phase has a very similar appearance and function to the Reference State of Community 1.1, except that it has a minor amount of cool-season exotic grasses and forbs. This phase functions at a high level for native wildlife; therefore, managers should consider the 2.0 Community Phase Pathways to avoid transitioning to State 3.0.

Invertebrates: Provides similar life requisites as Community Phase 1.1.

Birds: Provides similar life requisites as Community Phase 1.1.

Mammals: Provides similar life requisites as Community Phase 1.1.

Amphibians/Reptiles: Provides similar life requisites as Community Phase 1.1.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

Community Phase 2.2 Blue Grama/Sedges/Western Wheatgrass: Community Phase Pathway 2.1A is characterized by multi-year drought with or without heavy continuous grazing. Plant community diversity is reduced with a decline of deeper-rooted native species which are replaced by shorter, shallow-rooted grasses, and sedges. This plant community is on the cusp of crossing the threshold to the 3.0 Invaded State. Prescribed grazing with adequate recovery periods between grazing will shift the competitive edge to native species along Community Phase Pathway 2.2A; this is the most effective method to regain diverse cool-season grass and forb components in Community Phase 2.1. Every effort should be used to manage within Community Phase Pathway 2.2A to avoid crossing the threshold into State 3.0. Restoration Pathway R3A requires intensive management and economic inputs to successfully cross back to State 2.0.

Insects: Provides similar life requisites as Community Phase 1.2.

Birds: Provides similar life requisites as Community Phase 1.2

Mammals: Provides similar life requisites as Community Phase 2.2.

Amphibians/Reptiles: Provides similar life requisites as Community Phase 1.1.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

Community Phase 2.3 Needlegrasses/Exotic Cool-Season Grasses/Forbs: Long-term non-use or very light grazing, and no fire (along Community Pathway 2.1B or 2.2B) leads to a community phase characterized by shorter-statured grasses, including the increase of cool-season exotic grass and the reduction of prairie sandreed. This is an “at risk” plant community with high amounts of Kentucky bluegrass out-completing native grasses. Every effort should be made by managers to avoid implementing a grazing system that may favor reaching the 30 percent threshold for Kentucky bluegrass via Transitional Pathway T2A. When this plant community transitions over to State 3.0, it has little chance of restoration back to State 2.0 by implementing a prescribed grazing system and introducing prescribed fire.

Invertebrates: An increase exotic cool-season grass and a reduction in forbs reduces pollen and nectar sources, limiting use by pollination insect species. White heath aster, goldenrod, and white sagebrush are the main pollinator species. Increased litter may begin to limit ground nesting insect opportunities.

Birds: Grassland-nesting birds that favor short-statured vegetative cover will use this site. However, long-term non-use and no fire increases exotic cool-season exotic grasses, reducing use by grassland nesting birds.

Mammals: As exotic cool-season grasses increase, thermal, protective or escape cover becomes limited.

Amphibians and Reptiles: Provides similar life requisites as Community Phase 1.1.

Fish and Mussels: Although this ecological site is not typically adjacent to streams, rivers, or water bodies, runoff from Community Phase 2.3 increases due to management that increases short- statured grass and reduces bunch grasses. Management of this community phase, in conjunction with neighboring run-on sites, will have an indirect negative effect on aquatic species in streams and/or tributaries receiving water from Sands and adjacent sites.

3.0 Invaded State

Community Phase 3.1 Exotic Cool-Season Grasses/Forbs: Community Phase Pathway T2A generally occurs during long-term non-use or with heavy season-long grazing. This plant community phase is characterized by a dominance (more than 30 percent) to a complete dominance of exotic cool-season grasses (such as Kentucky bluegrass, crested wheatgrass, and smooth brome). Western snowberry tends to increase in density and cover. Restoration Pathway R3A requires remnant amounts of native warm-season grasses (i.e., blue grama), cool-season grasses (i.e., needlegrasses, western wheatgrass, prairie Junegrass), and forbs (i.e., silverleaf Indian breadroot, upright prairie coneflower). These remnant populations can only be expressed through frequent prescribed burns and high levels of prescribed grazing management targeting the exotic cool-season grasses. Intensified management along the R3A pathway will have significant short-term negative impacts on wildlife habitat; however, this is necessary to restore long-term native habitat functions.

Invertebrates: Non-use or low intensity (less than 20 percent utilization) grazing limits use by beneficial insects provided in States 1.0 and 2.0. Increased litter and lack of grazing leads to limited contact between plant material and mineral soil; this results in a cooler micro-climate which is unfavorable to most insects. Lack of bare soil limits ground nesting sites for native bees and other ground-nesting insects. The lack of nectar-producing plants limits forage opportunities for bumblebees, regal fritillary, monarch butterfly, and other pollinating species.

Birds: This homogeneous community phase, dominated by exotic plant species, provides limited habitat and life requisites for most obligate grassland-nesting birds. Lack of stature and plant diversity, along with increased litter and the tendency of Kentucky bluegrass and smooth brome to lay down, limits use by many grassland-nesting birds. Litter accumulations reduce use by chestnut-collared and McCown’s longspurs. Western snowberry reduces use of this site by species that avoid areas with woody vegetation. Sharp-tailed grouse may use these sites for brood rearing and winter cover; however, the reduction in forbs may limit foraging opportunities for chicks.

Mammals: Litter accumulation favors thermal, protective, and escape cover for small rodents. Large mammal thermal, protective, and escape cover becomes limited.

4.0 Go-Back State

Community Phase 4.1 Annual/Pioneer Perennial/Exotics: These plant communities are the result of severe soil disturbance (such as cropping, recreational activity, or concentrated livestock activity for a prolonged period). Following cessation of disturbances, the resulting plant community is dominated by early pioneer annual and perennial plant species. Plant species composition and production are highly variable. Weedy plants can provide pollinator habitat along with spring and summer cover for many mammals and birds, and their young. Dense weed cover can keep soils moist, increasing insect presence. Tall stature provided by some weeds, such as marsh elder and ragweed, offer thermal cover and seeds throughout winter.

Successful restoration of native species along Transition Pathway R4A results in a native grass and forb community in State 2.0. Failed restoration to native species through Restoration Pathway R4B results in Invaded State 3.0. Wildlife species response will be dependent upon plant community composition, vegetative structure, patch size, and management activities (such as prescribed grazing, burning, range planting, haying, or noxious weed control).

Animal Community – Grazing Interpretations

This site is well adapted to managed grazing by domestic livestock. The predominance of herbaceous plants across all plant community phases best lends these sites to grazing by cattle, but other domestic grazers with differing diet preferences may also be a consideration depending upon management objectives. Often, the current plant community does not match any particular plant community (as described in the ecological site description). Because of this, a resource inventory is necessary to document plant composition and production. Proper interpretation of this inventory data will permit the establishment of a safe, initial stocking rate for the type and class of animals and level of grazing management. More accurate stocking rate estimates should eventually be calculated using actual stocking rate information and monitoring data.

NRCS defines prescribed grazing as “managing the harvest of vegetation with grazing and/or browsing animals with the intent to achieve specific ecological, economic, and management objectives”. As used in this site description, the term ‘prescribed grazing’ is intended to include multiple grazing management systems (e.g., rotational grazing, twice-over grazing, conservation grazing, targeted grazing, etc.) provided that, whatever management system is implemented, it meets the intent of prescribed grazing definition.

The basic grazing prescription addresses balancing forage demand (quality and quantity) with available forage, varying grazing and deferment periods from year-to-year, matching recovery/deferment periods to growing conditions when pastures are grazed more than once in a growing season, implementation of a contingency (e.g., drought) plan, and a monitoring plan. When the management goal is to facilitate change from one plant community phase or state to another, then the prescription needs to be designed to shift the competitive advantage to favor the native grass and forb species.

Grazing levels are noted within the plant community narratives and pathways in reference to grazing/prescribed grazing management. “Degree of utilization” is defined as the proportion of the current years forage production that is consumed and/or destroyed by grazing animals (may refer to a single plant species or a portion or all the vegetation). “Grazing utilization” is classified as slight, moderate, full, close, and severe (see the following table for description of each grazing use category). The following utilization levels are also described in the Ranchers Guide to Grassland Management IV. Utilization levels are determined by using the landscape appearance method as outlined in the Interagency Technical Reference “Utilization Studies and Residual Measurements” 1734-3.

Utilization Level % Use Description
Slight (Light) 0-20 Appears practically undisturbed when viewed obliquely. Only choice areas and forage utilized.

Moderate 20-40 Almost all of accessible range shows grazing. Little or no use of poor forage. Little evidence of trailing to grazing.

Full 40-60 All fully accessible areas are grazed. The major sites have key forage species properly utilized (about half taken, half left). Points of concentration with overuse limited to 5 to 10 percent of accessible area.

Close (Heavy) 60-80 All accessible range plainly shows use and major sections closely cropped. Livestock forced to use less desirable forage, considering seasonal preference.

Severe > 80 Key forage species completely used. Low-value forages are dominant.

Hydrological functions

Water is the principal factor limiting forage production on this site. This site is dominated by soils in hydrologic group C. Infiltration varies from moderate to moderately rapid; runoff potential for this site varies from medium to high depending on surface texture, slope percent, and ground cover. The dense, claypan layer slows water movement through the soil profile. In many cases, areas with greater than 75% ground cover have the greatest potential for high infiltration and lower runoff. An example of an exception would be where shortgrasses form a strong sod and dominate the site. Dominance by blue grama, Kentucky bluegrass, and/or smooth brome will result in reduced infiltration and increased runoff. Areas where ground cover is less than 50% have the greatest potential to have reduced infiltration and higher runoff (refer to Section 4, NRCS National Engineering Handbook for runoff quantities and hydrologic curves).

Recreational uses

Hunting and Bird Watching: National wildlife refuges, waterfowl production areas, state wildlife management areas (WMA), and North Dakota and South Dakota Department of Trust Lands provide herbaceous and woody cover for wildlife. National Wildlife Refuges and waterfowl production areas are owned and managed by the United States Fish and Wildlife Service and are available for public hunting, hiking, and bird watching. In addition, the United States Army Corps of Engineers and the United States Bureau of Reclamation manage three man-made reservoirs - Jamestown Reservoir (2,036 acres), Pipestem Reservoir (1,027 acres), and Lake Ashtabula (5,174 acres) for flood control, also providing fish habitat and adjacent uplands for wildlife cover. Lonetree WMA is the largest state managed wildlife area covering 32,800 acres. Numerous WMAs in North Dakota and Game Production Areas in South Dakota are found within this MLRA. The largest refuges managed by the United States Fish and Wildlife service are Arrowwood National Wildlife Refuge (NWR) Complex consists of 75,000 acres and Tewaukon National NWR covers 8,363 acres.

Fishing: Approximately 100 lakes are managed for public fishing within MLRA 55B. Most of these lakes offer boat docks and ramps. These lakes contain various sport fish including walleye, northern pike, yellow perch, catfish, trout, crappie, and bluegill. Many of these lakes are known for excellent round-around walleye and yellow perch fishery.

Camping: Fort Ramson State Park, Pipestem Reservoir, Jamestown Reservoir, Spiritwood Lake, Clausen Springs, Little Yellowstone, Richmond Lake State Recreation Area, Mina Lake State Recreation Area, and other public and private campgrounds are found within the MLRA. Limited, primitive camping is available on wildlife management areas. Ft. Ransom State Park (North Dakota), located along the Sheyenne River has a designated horse park with 15 miles of trails.

Hiking/Biking/Horseback Riding: Horseback riders, hikers, and biker can enjoy over 15 miles of multi-use trails at Fort Ransom State Park. The Jamestown Reservoir (5 miles), Pipestem Reservoir (8 miles) and Arrowwood National Wildlife Refuge (9.4 miles) maintain hiking trails. The Lonetree Wildlife Management Area has a 32- mile segment of the North Country Trail. It is designed for hiking and non-motorized travel including mountain bikes or horseback riding.

Canoeing/Kayaking: The Sheyenne River offers 278 miles of canoeing/kayaking from May-July. A kayak kiosk is located at Valley City and canoe/kayak rentals are available at Fort Ransom State Park. The James River has a canoe trail starting in Grand Rapids and canoeing down to the James River Dam site in LaMoure; no rentals are available.

Auto Tour: A 63-mile scenic drive starts north of Valley City and heading south through Sheyenne River Valley. Audubon National Wildlife Refuge offers a 5.5-mile auto-tour route winding through both prairie grassland and wetland habitats of the lower portion of the James River Valley.

Wood products

There are no significant wood products found on this site.

Other products

Seed harvest of native plant species can provide additional income on this site.

Other information

Site Development and Testing Plan
• Further evaluation is needed on the drainage class/seasonal high water table. The Letcher OSD allows somewhat poorly drained phases. Current soils correlations are almost entirely moderately well drained phases. Letcher minor components commonly occur in map units with predominantly poorly drained soils (e,g, Stirum). On-site investigation is needed to determine if plant communities on the somewhat poorly drained phase of Letcher better fits Sandy Claypan, Saline Subirrigated, or Saline Lowland ESD.
• Further evaluation and refinement of the State-and-Transition model may be needed to identify disturbance driven dynamics. Additional states and/or phases may be required to address grazing response.
• Further documentation may be needed for plant communities in all states. Plant data has been collected in previous range-site investigations, including clipping data; however, this data needs review. If geo-referenced sites meeting Tier 3 standards for either vegetative or soil data are not available, representative sites will be selected for further investigation.
• Site concepts will be refined as the above noted investigations are completed.
• The long-term goal is to complete an approved, correlated Ecological Site Description as defined by the National Ecological Site Handbook.

This ESD is the best available knowledge. The site concept and species composition table have been used in the field and tested for more than five years. It is expected that as additional information becomes available revisions may be required.

Inventory data references

Information presented here has been derived from NRCS and other federal/state agency clipping and inventory data. Also, field knowledge of range-trained personnel was used. All descriptions were peer reviewed and/or field-tested by various private, state and federal agency specialists.

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Contributors

Jeff Printz, NRCS State Range Management Specialist, Retired
Stan Boltz, NRCS Range Health Specialist
Jody Forman, NRCS Range Management Specialist
David Dewald, NRCS State Biologist, Retired
Mark Hayek, State Range Management Specialist, Retired
Kevin Sedivec, Extension Range Managements Specialist
Shawn Dekeyser, North Dakota State University
Rob Self, The Nature Conservancy
Lee Voight, NRCS Range Management Specialist

Approval

Suzanne Mayne-Kinney, 4/25/2025